Glycoside Hydrolase Family 5

Activities in Familyendo-β-1,4-glucanase / cellulase (EC 3.2.1.4); endo-β-1,4-xylanase (EC 3.2.1.8); β-glucosidase (EC 3.2.1.21); β-mannosidase (EC 3.2.1.25); β-glucosylceramidase (EC 3.2.1.45); glucan β-1,3-glucosidase (EC 3.2.1.58); licheninase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); glucan endo-1,6-β-glucosidase (EC 3.2.1.75); mannan endo-β-1,4-mannosidase (EC 3.2.1.78); cellulose β-1,4-cellobiosidase (EC 3.2.1.91); steryl β-glucosidase (EC 3.2.1.104); endoglycoceramidase (EC 3.2.1.123); chitosanase (EC 3.2.1.132); β-primeverosidase (EC 3.2.1.149); xyloglucan-specific endo-β-1,4-glucanase (EC 3.2.1.151); endo-β-1,6-galactanase (EC 3.2.1.164); hesperidin 6-O-α-L-rhamnosyl-β-glucosidase (EC 3.2.1.168); β-1,3-mannanase (EC 3.2.1.-); arabinoxylan-specific endo-β-1,4-xylanase (EC 3.2.1.-); mannan transglycosylase (EC 2.4.1.-)
Mechanism Retaining
ClanGH-A
3D Structure Status( β / α ) 8
Catalytic Nucleophile/BaseGlu (experimental)
Catalytic Proton DonorGlu (experimental)
NoteOnce known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)MEGAZYME; NZYTech; PROZOMIX;
Statistics GenBank accession (12399); Uniprot accession (1922); PDB accession (212); 3D entries (75); cryst (0)
All (11020) Archaea (67) Bacteria (8762) Eukaryota (2065) Viruses (6) unclassified (120) Structure (75) Characterized (547)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | ... | 88 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D Subf
 BAMY6639_00620   Bacillus amyloliquefaciens UMAF6639 AMQ71229.1     2
 endoglucanase A (EngA)   Bacillus amyloliquefaciens UMAS1002 AAL99668.1 Q8RPQ6   2
 BSO20_02700   Bacillus amyloliquefaciens WS-8 APQ49018.1     2
 BAMY_08970   Bacillus amyloliquefaciens Y14 APB82271.1     2
 MUS_2161 (BglC)   Bacillus amyloliquefaciens Y2 AFJ62120.1     2
 BaGK_15910 (fragment)   Bacillus atrophaeus GQJK17 ASS72344.1     2
 D068_cds30840 (fragment)   Bacillus atrophaeus UCMB-5137 AKL85635.1     2
 CY96_16600   Bacillus bombysepticus str. Wang AHX19543.1     48
 Bcell_1191   Bacillus cellulosilyticus DSM 2522 ADU29456.1 D2LUH5   4
 Bcell_3371   Bacillus cellulosilyticus DSM 2522 ADU31613.1 D2LQZ8   2
 endo-β-1,4-glucanase A (CelA;EgA;Bcell_0438) 3.2.1.4 Bacillus cellulosilyticus DSM 2522 ADU28720.1
AAA22301.1
D2LVP1
P06566
  2
 endo-β-1,4-glucanase B (CelB;EgB;Bcell_0437) 3.2.1.4 Bacillus cellulosilyticus DSM 2522 ADU28719.1
AAA22299.1
D2LVP0   2
 endo-β-1,4-glucanase C (CelC;EgC;Cel2;Bcell_3370) 3.2.1.4 Bacillus cellulosilyticus DSM 2522 ADU31612.1
AAA22306.1
D2LQZ9
P19570
  2
 B5E38_0519   Bacillus cereus 25 ARO58138.1     48
 DA68_13320   Bacillus cereus A1 ANE86586.1     48
 BCB4264_A3539   Bacillus cereus B4264 ACK60962.1 B7H8S5   48
 WR51_17990   Bacillus cereus CMCC P0011 ANC14724.1     48
 WR47_17990   Bacillus cereus CMCC P0021 ANC08907.1     48
 BA203_16885   Bacillus cereus D12_2 ASI75731.1     48
 FORC5_3317   Bacillus cereus FORC_005 AKE17854.1     48
 FORC13_4232   Bacillus cereus FORC_013 ALZ63293.1     48
 FORC47_3413   Bacillus cereus FORC_047 ASL66258.1     48
 FORC48_3449   Bacillus cereus FORC_048 ASI84531.1     48
 BCG9842_B1694   Bacillus cereus G9842 ACK97616.1 B7IRG0   48
 endo-β-1,4-glucanase (fragment)   Bacillus cereus IARI-SP-7 AGW99984.1     2
 BA204_16740   Bacillus cereus K8 ASJ49621.1     48
 BA201_17330   Bacillus cereus M13 ASK17637.1     48
 C1T25_10255   Bacillus cereus MBGJa3 AUZ28685.1     2
 CSW12_10225   Bacillus cereus MLY1 AUB63409.1     48
 ACN91_10355   Bacillus cereus NJ-W ALC55172.1     48
 cellulase   Bacillus cereus Y5 AGK07581.1     48
 β-mannanase   Bacillus circulans AAS48170.1 Q6QHT4   8
 β-mannanase (TM1) 3.2.1.78 Bacillus circulans AAT39478.1 Q6IUP2   8
 β-mannanase (Man1;ManB48) 3.2.1.78 Bacillus circulans CGMCC 1416 AAX87002.1 Q49HI4   8
 β-mannanase (Man5A) 3.2.1.78 Bacillus circulans CGMCC1554 AAX87003.1 Q0IJR6   8
 β-mannanase (G) 3.2.1.78 Bacillus circulans K-1 BAA25878.1 O66185   8
 BC8716_04830   Bacillus clausii DSM 8716 AST98531.1     8
 DB29_01746   Bacillus clausii ENTPro ALA52574.1     8
 ABC0320   Bacillus clausii KSM-K16 BAD62862.1 Q5WL93   8
 BGM20_03610   Bacillus gibsonii FJAT-10019 AOL29768.1     2
 BGLY_1877   Bacillus glycinifermentans SCA85700.1     7
 BGLY_1876   Bacillus glycinifermentans SCA85699.1     4
 BGLY_2367 (Bglc2)   Bacillus glycinifermentans SCA86190.1     2
 COP00_03840   Bacillus glycinifermentans KBN06P03352 ATH91857.1     2
 β-glucanase (CelB;BhCel5B;BH0603) (Cel5B) 3.2.1.4 Bacillus halodurans C-125 BAB04322.1
NP_241469.1
Q9KF82 4V2X[A] 4
 Cellulase (fragment)   Bacillus halodurans TSPV1 ALL28250.1     4
 BleG1_0468   Bacillus lehensis G1 AIC93076.1     54
 endo-β-1,4-glucanase, partial   Bacillus licheniformis AIY72754.1     2
 endo-β-1,4-glucanase, partial   Bacillus licheniformis AIY72755.1     2
 endo-β-1,4-glucanase, partial   Bacillus licheniformis AIY72756.1     2
 endo-β-1,4-glucanase (Egl01) 3.2.1.4 Bacillus licheniformis AIY72753.1     2
 CPQ91_09265   Bacillus licheniformis ATCC 9789 ATI76022.1     7
 CPQ91_09260   Bacillus licheniformis ATCC 9789 ATI76021.1     4
 CPQ91_10290   Bacillus licheniformis ATCC 9789 ATI78443.1     2
 BL1202_02034   Bacillus licheniformis BL1202 AOP14982.1     4
 BL1202_02035   Bacillus licheniformis BL1202 AOP14983.1     7
 BL1202_02293   Bacillus licheniformis BL1202 AOP15241.1     2
 BLi01883 / BL01229   Bacillus licheniformis DSM 13 = ATCC 14580 AAU40778.1
AAU23418.1
Q65JI6   7
 endoglucanase (BlCel5B;BLi01882;BL01230) (Cel5B) 3.2.1.4 Bacillus licheniformis DSM 13 = ATCC 14580 AAU40777.1
AAU23417.1
Q65JI7 4YZP[A]
4YZT[A]
4
 endo-β-1,4-glucanase C (BglC;BLi02088;BL01471) (Cel5A) 3.2.1.4 Bacillus licheniformis DSM 13 = ATCC 14580 AAU23613.2
AAU23613.1
AAU40977.1
ACR87895.1
CAF05574.1
CAV28462.1
A5A671
Q65IY7
  2
 endo-β-1,4-glucanase (CelA)   Bacillus licheniformis F11 CAJ70714.1 Q1EM84   2
 CelB   Bacillus licheniformis F11 CAJ70720.1 Q5K2M0   4
 endo-β-1,4-glucanase (CelA)   Bacillus licheniformis F5 CAJ70710.1 Q1EM84   2
 CelB   Bacillus licheniformis F5 CAJ70717.1 Q5K2M0   4
 endo-β-1,4-glucanase (Cel5A) (Cel5A) 3.2.1.4 Bacillus licheniformis GXN151 / B-41361 AAP51020.1 Q7X3S6   2
 AB684_08870   Bacillus licheniformis HRBL-15TDI7 AMR10287.1     4
 AB684_09875   Bacillus licheniformis HRBL-15TDI7 AMR12722.1     2
 AB684_08875   Bacillus licheniformis HRBL-15TDI7 AMR10288.1     7
 endo-β-1,4-glucanase   Bacillus licheniformis IARI-SP-3 AGW99979.1     2
 cellulase   Bacillus licheniformis K11 ABK63476.1 A0T1G5   2
 C1T27_09545   Bacillus licheniformis MBGJa67 AUZ30578.1     7
 C1T27_09540   Bacillus licheniformis MBGJa67 AUZ30577.1     4
 C1T27_10580   Bacillus licheniformis MBGJa67 AUZ32937.1     2
 endoglucanase (CelA) 3.2.1.4 Bacillus licheniformis MD1 CAE82178.1 Q5QSM2   2
 CelB   Bacillus licheniformis MD1 CAH10344.1 Q5K2M0   4
 YdhT (fragment)   Bacillus licheniformis MD1 CAH10345.1 Q5K2L9   7
 cellulose hydrolase (Cel5H)   Bacillus licheniformis MSB03 ACY72379.1 D1L8C4   2
 Cel5C (Cel5C)   Bacillus licheniformis MSB03 ACY72382.1 D1L8C7   4
 Cel5D (Cel5D)   Bacillus licheniformis MSB03 ACY72383.1 D1L8C8   7
 B37_01402   Bacillus licheniformis SCCB 37 ARC73454.1     7
 B37_01607 (EglS)   Bacillus licheniformis SCCB 37 ARC73659.1     2
 B37_01401 (EngD)   Bacillus licheniformis SCCB 37 ARC73453.1     4
 B14_02580   Bacillus licheniformis SCDB 14 ARC65578.1     7
 B14_02581 (EngD)   Bacillus licheniformis SCDB 14 ARC65579.1     4
 B14_02377 (EglS)   Bacillus licheniformis SCDB 14 ARC65375.1     2
 B34_00221 (EngD)   Bacillus licheniformis SCDB 34 ARC67664.1     4
 B34_00630 (EglS)   Bacillus licheniformis SCDB 34 ARC68073.1     2
 B34_00222   Bacillus licheniformis SCDB 34 ARC67665.1     7
 BaDB11_04388 (EngD)   Bacillus licheniformis SCK B11 ARC62951.1     4
 BaDB11_04389   Bacillus licheniformis SCK B11 ARC62952.1     7
 BaDB11_00288 (EglS)   Bacillus licheniformis SCK B11 ARC58957.1     2
 S100141_01268   Bacillus licheniformis SRCM100141 ARW42590.1     4
 S100141_01269   Bacillus licheniformis SRCM100141 ARW42591.1     7
 S100141_01476   Bacillus licheniformis SRCM100141 ARW42798.1     2
 S101441_02103   Bacillus licheniformis SRCM101441 ARW31652.1     2
 glycoside hydrolase (Gh5C)   Bacillus licheniformis SVD1 BAL45506.1     7
 glycoside hydrolase (Gh5B)   Bacillus licheniformis SVD1 BAL45505.1     4
 endoglucanase (Gh5A)   Bacillus licheniformis SVD1 BAL45504.1     2
 MUY_001893 (CelC)   Bacillus licheniformis WX-02 AKQ73025.1     4
 MUY_002122 (BglC)   Bacillus licheniformis WX-02 AKQ73254.1     2
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