Glycoside Hydrolase Family 5

Activities in Familyendo-β-1,4-glucanase / cellulase (EC 3.2.1.4); endo-β-1,4-xylanase (EC 3.2.1.8); β-glucosidase (EC 3.2.1.21); β-mannosidase (EC 3.2.1.25); β-glucosylceramidase (EC 3.2.1.45); glucan β-1,3-glucosidase (EC 3.2.1.58); licheninase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); glucan endo-1,6-β-glucosidase (EC 3.2.1.75); mannan endo-β-1,4-mannosidase (EC 3.2.1.78); cellulose β-1,4-cellobiosidase (EC 3.2.1.91); steryl β-glucosidase (EC 3.2.1.104); endoglycoceramidase (EC 3.2.1.123); chitosanase (EC 3.2.1.132); β-primeverosidase (EC 3.2.1.149); xyloglucan-specific endo-β-1,4-glucanase (EC 3.2.1.151); endo-β-1,6-galactanase (EC 3.2.1.164); hesperidin 6-O-α-L-rhamnosyl-β-glucosidase (EC 3.2.1.168); β-1,3-mannanase (EC 3.2.1.-); arabinoxylan-specific endo-β-1,4-xylanase (EC 3.2.1.-); mannan transglycosylase (EC 2.4.1.-)
Mechanism Retaining
ClanGH-A
3D Structure Status( β / α ) 8
Catalytic Nucleophile/BaseGlu (experimental)
Catalytic Proton DonorGlu (experimental)
NoteOnce known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)MEGAZYME; NZYTech; PROZOMIX;
Statistics GenBank accession (11690); Uniprot accession (1922); PDB accession (209); 3D entries (74); cryst (0)
All (10313) Archaea (67) Bacteria (8119) Eukaryota (2005) Viruses (5) unclassified (117) Structure (74) Characterized (547)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | ... | 82 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D Subf
 AORI_3215   Amycolatopsis orientalis HCCB10007 AGM05800.1     51
 AORI_3875   Amycolatopsis orientalis HCCB10007 AGM06460.1     2
 AORI_5568   Amycolatopsis orientalis HCCB10007 AGM08151.1     2
 endo-β-1,4-glucanase (End2)   Anabaena laxa RPAN8 / CCC 381 ADE22239.1 D5KTA0  
 ANA_C11786   Anabaena sp. 90 AFW94547.1    
 ANT_03660   Anaerolinea thermophila UNI-1 BAJ62400.1     37
 ANT_22130   Anaerolinea thermophila UNI-1 BAJ64239.1     12
 ANT_18750   Anaerolinea thermophila UNI-1 BAJ63901.1     25
 BEQ56_08495   Anaerolineaceae bacterium oral taxon 439 W11661 AOH43511.1     4
 Anae109_4201   Anaeromyxobacter sp. Fw109-5 ABS28379.1 A7HI33  
 Anae109_3225   Anaeromyxobacter sp. Fw109-5 ABS27420.1 A7HFC4  
 GFC28_925   Anoxybacillus sp. B2M1 ANB56319.1     22
 GFC29_701   Anoxybacillus sp. B7M1 ANB65284.1     22
 SAMN00777080_1515   Aquiflexum balticum DSM 16537 SMD42944.1     25
 SAMN00777080_2486   Aquiflexum balticum DSM 16537 SMD43874.1     38
 SAMN00777080_1055   Aquiflexum balticum DSM 16537 SMD42501.1     36
 A9P82_12630   Arachidicoccus sp. BS20 ANI90055.1     13
 A9P82_13735   Arachidicoccus sp. BS20 ANI90259.1     2
 Arch_0033   Arcanobacterium haemolyticum DSM 20595 ADH91800.1 D7BLK1   18
 SAMN04489737_0425   Arcanobacterium phocae DSM 10002 SDU78319.1     18
 SAMN04489737_0244   Arcanobacterium phocae DSM 10002 SDU77936.1     28
 AA314_01832   Archangium gephyra DSM 2261 AKJ00206.1     7
 AREALGSMS7_03094   Arenibacter algicola SMS7 ASO06525.1     13
 CCB80_07970   Armatimonadetes bacterium Uphvl-Ar1 ARU41080.1     47
 AOC05_14730   Arthrobacter alpinus R3.8 ALE93292.1     18
 CGQ24_15805   Arthrobacter sp. 7749 ASN40324.1     18
 MB46_17100   Arthrobacter sp. A3 ALV46941.1     18
 UM93_09880   Arthrobacter sp. IHBB 11108 AJT41747.1     18
 ART_2884   Arthrobacter sp. PAMC 25486 AIY02483.1     18
 ART_1927   Arthrobacter sp. PAMC 25486 AIY01526.1    
 CGK93_22015   Arthrobacter sp. YN ASN22040.1     18
 CGK93_22205   Arthrobacter sp. YN ASN22069.1     18
 Astex_2349   Asticcacaulis excentricus CB 48 ADU14002.1 C5SHS4   48
 Astex_2497   Asticcacaulis excentricus CB 48 ADU14148.1 C5SQ05   25
 Astex_2880   Asticcacaulis excentricus CB 48 ADU14517.1 C5SNP2   7
 Astex_3212   Asticcacaulis excentricus CB 48 ADU14847.1 C5SP55   4
 Apar_1088   Atopobium parvulum DSM 20469 ACV51516.1 C8W7S5  
 M673_19220   Aureimonas sp. AU20 ALN74859.1    
 dqs_2441   Azoarcus olearius DQS4 ANQ85471.1     45
 dqs_2369   Azoarcus olearius DQS4 ANQ85400.1     5
 endoglucanase (Egl;EglA;azo2236) 3.2.1.4 Azoarcus sp. BH72 ABB92850.1
CAL94853.1
A0S6A6   5
 azo2311   Azoarcus sp. BH72 CAL94928.1 A1K7X3   45
 AzCIB_4213   Azoarcus sp. CIB AKU14106.1     45
 AZKH_4274 (Spr1)   Azoarcus sp. KH32C BAL26553.1     45
 endo-β-1,4-glucanase (Egl;AZC_1301) 3.2.1.4 Azorhizobium caulinodans ORS 571 BAF87299.1 A8I0V5   1
 ABAZ39_32285   Azospirillum brasilense Az39 AIB16520.1     1
 ABAZ39_20660   Azospirillum brasilense Az39 AIB14334.1     1
 ABAZ39_04560   Azospirillum brasilense Az39 AIB11296.1     1
 AMK58_20535   Azospirillum brasilense Sp 7 ALJ37806.1     1
 AMK58_26700   Azospirillum brasilense Sp 7 ALJ39073.1     1
 AMK58_09480   Azospirillum brasilense Sp 7 ALJ35637.1     1
 AZOBR_150049   Azospirillum brasilense Sp245 CCC98631.1     1
 AZOBR_p1110164   Azospirillum brasilense Sp245 CCD00695.1     1
 AZOBR_p470008   Azospirillum brasilense Sp245 CCD03592.1     1
 AZOLI_p10561   Azospirillum lipoferum 4B CBS88800.1     1
 AZOLI_p40099   Azospirillum lipoferum 4B CBS90502.1     1
 AZL_a06890   Azospirillum sp. B510 BAI74220.1 D3P0P1   1
 AZL_d05040   Azospirillum sp. B510 BAI76330.1 D3P6Q1   1
 AL072_19540   Azospirillum thiophilum BV-S ALG73272.1     1
 AL072_23195   Azospirillum thiophilum BV-S ALG73945.1     1
 ACG10_17745   Azotobacter chroococcum B3 ASL27949.1     45
 Achr_34060   Azotobacter chroococcum NCIMB 8003 AJE22811.1     45
 AvCA_07130   Azotobacter vinelandii CA AGK15534.1     45
 AvCA6_07130   Azotobacter vinelandii CA6 AGK19467.1     45
 Avin_07130   Azotobacter vinelandii DJ ATCC BAA-1303 ACO76963.1 C1DLL5   45
 endo-β-1,4-glucanase (Egl-1139) / III-3A (Cel5A) 3.2.1.4 Bacillus akibai / KSM-1139 BAA00045.1
AAA22305.1
ACE14924.1
P06564   2
 endoglucanase (EG-I-1)   Bacillus akibai I-1 AFI25187.1     2
 CSE15_08310   Bacillus altitudinis P-10 ATP93954.1     7
 CFN77_08490   Bacillus altitudinis SGAir0031 ATH72233.1     7
 endoglucanase   Bacillus amyloliquefaciens AMS1 AKS29707.1     2
 A1R12_09055   Bacillus amyloliquefaciens B15 AMR50503.1     2
 endo-β-1,4-glucanase (EgL)   Bacillus amyloliquefaciens BY-5 AMW90767.1     2
 U471_18630   Bacillus amyloliquefaciens CC178 AGZ56565.1     2
 endo-β-1,4-glucanase   Bacillus amyloliquefaciens IARI-SP-2 AGW99978.1     2
 KSO_010315   Bacillus amyloliquefaciens IT-45 AGF27558.1     2
 KHU1_1621   Bacillus amyloliquefaciens KHG19 AJK65578.1     2
 SB45_09085   Bacillus amyloliquefaciens L-H15 AJH24169.1     2
 XM40_09090   Bacillus amyloliquefaciens L-S60 AKD22364.1     2
 U722_09575   Bacillus amyloliquefaciens LFB112 AHC42348.1     2
 BSF20_16750   Bacillus amyloliquefaciens LM2303 APH49957.1     2
 cellulase (fragment)   Bacillus amyloliquefaciens MBAA3 AHJ80840.1     2
 AVM03_03620   Bacillus amyloliquefaciens MBE1283 ALV01512.1     2
 endoglucanase (EglII) 3.2.1.4 Bacillus amyloliquefaciens PSM 3.1 ADC54852.1 D3Y2F0   2
 AS588_06260   Bacillus amyloliquefaciens S499 AMP31635.1     2
 BAMY6614_13370   Bacillus amyloliquefaciens UMAF6614 AMQ74283.1     2
 BAMY6639_00620   Bacillus amyloliquefaciens UMAF6639 AMQ71229.1     2
 endoglucanase A (EngA)   Bacillus amyloliquefaciens UMAS1002 AAL99668.1 Q8RPQ6   2
 BSO20_02700   Bacillus amyloliquefaciens WS-8 APQ49018.1     2
 BAMY_08970   Bacillus amyloliquefaciens Y14 APB82271.1     2
 MUS_2161 (BglC)   Bacillus amyloliquefaciens Y2 AFJ62120.1     2
 BaGK_15910 (fragment)   Bacillus atrophaeus GQJK17 ASS72344.1     2
 D068_cds30840 (fragment)   Bacillus atrophaeus UCMB-5137 AKL85635.1     2
 CY96_16600   Bacillus bombysepticus str. Wang AHX19543.1     48
 Bcell_1191   Bacillus cellulosilyticus DSM 2522 ADU29456.1 D2LUH5   4
 Bcell_3371   Bacillus cellulosilyticus DSM 2522 ADU31613.1 D2LQZ8   2
 endo-β-1,4-glucanase A (CelA;EgA;Bcell_0438) 3.2.1.4 Bacillus cellulosilyticus DSM 2522 ADU28720.1
AAA22301.1
D2LVP1
P06566
  2
 endo-β-1,4-glucanase B (CelB;EgB;Bcell_0437) 3.2.1.4 Bacillus cellulosilyticus DSM 2522 ADU28719.1
AAA22299.1
D2LVP0   2
 endo-β-1,4-glucanase C (CelC;EgC;Cel2;Bcell_3370) 3.2.1.4 Bacillus cellulosilyticus DSM 2522 ADU31612.1
AAA22306.1
D2LQZ9
P19570
  2
 B5E38_0519   Bacillus cereus 25 ARO58138.1     48
 DA68_13320   Bacillus cereus A1 ANE86586.1     48
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