Glycoside Hydrolase Family 5 / Subf 2

Activities in Familyendo-β-1,4-glucanase / cellulase (EC 3.2.1.4); endo-β-1,4-xylanase (EC 3.2.1.8); β-glucosidase (EC 3.2.1.21); β-mannosidase (EC 3.2.1.25); β-glucosylceramidase (EC 3.2.1.45); glucan β-1,3-glucosidase (EC 3.2.1.58); licheninase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); glucan endo-1,6-β-glucosidase (EC 3.2.1.75); mannan endo-β-1,4-mannosidase (EC 3.2.1.78); cellulose β-1,4-cellobiosidase (EC 3.2.1.91); steryl β-glucosidase (EC 3.2.1.104); endoglycoceramidase (EC 3.2.1.123); chitosanase (EC 3.2.1.132); β-primeverosidase (EC 3.2.1.149); xyloglucan-specific endo-β-1,4-glucanase (EC 3.2.1.151); endo-β-1,6-galactanase (EC 3.2.1.164); hesperidin 6-O-α-L-rhamnosyl-β-glucosidase (EC 3.2.1.168); β-1,3-mannanase (EC 3.2.1.-); arabinoxylan-specific endo-β-1,4-xylanase (EC 3.2.1.-); mannan transglycosylase (EC 2.4.1.-)
Activities in Sub Family
Mechanism Retaining
ClanGH-A
3D Structure Status( β / α ) 8
Catalytic Nucleophile/BaseGlu (experimental)
Catalytic Proton DonorGlu (experimental)
NoteOnce known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)MEGAZYME; NZYTech; PROZOMIX;
Statistics GenBank accession (1085); Uniprot accession (275); PDB accession (42); 3D entries (14); cryst (0)
All (968) Bacteria (799) Eukaryota (155) unclassified (14) Structure (14) Characterized (119)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 |
Bacteria
Protein Name EC#Organism GenBankUniprotPDB/3DSubf
 CLOSACC_09830 (Cel5A)   Clostridium saccharobutylicum NCP 195 AQS13168.1     2
 CLOSC_09760 (Cel5A)   Clostridium saccharobutylicum NCP 200 AQR89279.1     2
 CLOBY_10270 (Cel5A)   Clostridium saccharobutylicum NCP 258 AQS08912.1     2
 endo-β-1,4-glucanase (EglA) 3.2.1.4 Clostridium saccharobutylicum P262 AAA23230.1 P15704   2
 Clo1100_1353   Clostridium sp. BNL1100 AEY65590.1     2
 CCU_20020   Coprococcus sp. ART55/1 CBK83458.1 D5HEX5   2
 CCU_25500   Coprococcus sp. ART55/1 CBK83877.1 D5HG44   2
 CYFUS_008414   Cystobacter fuscus DSM 52655 ATB42935.1     2
 CYFUS_001603   Cystobacter fuscus DSM 52655 ATB36189.1     2
 endoglucanase 5A (ChCel5A;CHU_1107) (Cel5A) 3.2.1.4 Cytophaga hutchinsonii ATCC 33406 ABG58383.1 Q11W33   2
 periplasmic endoglucanase 5B (CHU_2103) (Cel5B) 3.2.1.4 Cytophaga hutchinsonii ATCC 33406 ABG59366.1 Q11T98 5IHS[A] 2
 SAMN06298216_2068   Cytophagales bacterium TFI 002 SOE21610.1     2
 DTL3_1188   Defluviitoga tunisiensis CEP78490.1     2
 endocellulase (Cel5)   Deinococcus cellulosilyticus 5516J-15 AKN19928.1     2
 Dd1591_1529   Dickeya chrysanthemi Ech1591 ACT06384.1 C6CF34   2
 endo-β-1,4-glucanase (Cel5Z) 3.2.1.4 Dickeya chrysanthemi PY35 AAF18152.1 Q9REW0   2
 endo-β-1,4-glucanase Z (CelZ;EgZ;Dda3937_02793) (Cel5A=Cel5Z) 3.2.1.4 Dickeya dadantii 3937 ADM99099.1
CAA68604.1
P07103 1EGZ[A,B,C] 2
 DDI_2600   Dickeya dianthicola RNS04.9 ATO33768.1     2
 DDI_0695   Dickeya dianthicola RNS04.9 ATO31863.1     2
 CVE23_14355   Dickeya fangzhongdai DSM 101947 ATZ95058.1     2
 Dd703_1886   Dickeya paradisiaca Ech703 ACS85678.1 C6C568   2
 BJD21_19575   Dickeya solani D s0432-1 AUH10464.1     2
 A4U42_01465   Dickeya solani IPO 2222 ANE74123.1     2
 BJJ98_19545   Dickeya solani PPO 9019 AUH14400.1     2
 D083_0932   Dickeya solani RNS 08.23.3.1.A AUC41282.1     2
 W909_12595   Dickeya zeae EC1 AJC66868.1     2
 Dd586_1489   Dickeya zeae Ech586 ACZ76357.1 D2BWX3   2
 C1O30_13580   Dickeya zeae MS2 AUQ26034.1     2
 Echvi_1632   Echinicola vietnamensis DSM 17526 AGA77897.1     2
 EUBELI_20587   [Eubacterium] eligens ATCC 27750 ACR73731.1 C4Z6Y9   2
 EUBREC_0616   [Eubacterium] rectale ATCC 33656 ACR74405.1 C4ZCQ0   2
 EUR_02280   [Eubacterium] rectale DSM 17629 CBK89462.1 D6E222   2
 ERE_23730   [Eubacterium] rectale M104/1 CBK94241.1 D4JKU4   2
 EUS_00270   [Eubacterium] siraeum 70/3 CBK95395.1 D4JQM6   2
 ES1_04480   [Eubacterium] siraeum V10Sc8a CBL33590.1 D4MIK7   2
 AALO17_05750 (fragment)   Faecalibaculum rodentium Alo17 AMK53709.1     2
 AALO17_05740 (fragment)   Faecalibaculum rodentium Alo17 AMK53708.1     2
 Fisuc_1224 / FSU_1685 3.2.1.4 Fibrobacter succinogenes subsp. succinogenes S85 ACX74827.1
ADL26286.1
C9RQJ1   2
 Fisuc_1661 / FSU_2150   Fibrobacter succinogenes subsp. succinogenes S85 ACX75256.1
ADL25349.1
C9RRS0   2
 Fisuc_2011 / FSU_2534   Fibrobacter succinogenes subsp. succinogenes S85 ACX75598.1
ADL25634.1
C9RJ81   2
 endoglucanase (partial) 3.2.1.4 Fibrobacter succinogenes subsp. succinogenes S85 AAC06197.1 O66065   2
 cellulase (Fisuc_2364;FSU_2914) (Cel5H) 3.2.1.4 Fibrobacter succinogenes subsp. succinogenes S85 ABU45500.1
ACX75950.1
ADL27061.1
A7UG69   2
 FLA_2145   Filimonas lacunae NBRC 104114 BAV06130.1     2
 MY04_0191 (Cel5B)   Flammeovirga sp. MY04 ANQ47573.1     2
 D770_26615   Flammeovirgaceae bacterium 311 AHM63566.1     2
 D770_06795   Flammeovirgaceae bacterium 311 AHM59622.1     2
 BB050_02617 (EglS)   Flavobacterium johnsoniae GSE09 AOC95714.1     2
 Fjoh_4946   Flavobacterium johnsoniae UW101 ABQ07945.1 A5FA24   2
 ALW18_03010   Flavobacterium psychrophilum Z1 AOE54289.1     2
 AMR72_03010   Flavobacterium psychrophilum Z2 ALM50669.1     2
 endo-β-1,4-glucanase (CelA;GsCelA) 3.2.1.4 Geobacillus sp. 70PC53 ACJ60856.1 C5H6X3 4XZB[A]
4XZW[A]
2
 endoglucanase   Geobacillus stearothermophilus CICC-20649 ADO21451.1 E3V048   2
 endoglucanase (Cel5A)   Gynuella sunshinyii YC6258 AII80614.1     2
 YC6258_05810   Gynuella sunshinyii YC6258 AJQ97838.1     2
 YC6258_02995   Gynuella sunshinyii YC6258 AJQ95033.1     2
 endoglucanase (Cel5;HCH_03914) 3.2.1.4 Hahella chejuensis KCTC 2396 ABC30636.1 Q2SFD8   2
 SD1D_0239 (EglA)   Herbinix luporum SD1D CUH91792.1     2
 HVS_14300 (fragment)   Herbivorax saccincola GGR1 AUG58721.1     2
 HVS_12420 (Cel5A)   Herbivorax saccincola GGR1 AUG58357.1     2
 AXW84_14545   Hymenobacter sp. PAMC 26628 AMJ66514.1     2
 Hsw_1037   Hymenobacter swuensis DY53 AHJ96632.1     2
 BN2497_11653   Janthinobacterium sp. CG23_2 CUI24051.1     2
 AOZ06_10145   Kibdelosporangium phytohabitans KLBMP1111 ALG07233.1     2
 ORF   Kibdelosporangium sp. MJ126-NF4 CTQ95794.1     2
 B7C62_30465   Kitasatospora albolonga YIM 101047 ARF76115.1     2
 Kfla_1714   Kribbella flavida DSM 17836 ADB30809.1 D2PNF7   2
 cellulase, partial (Cel1) (fragment)   Lactobacillus plantarum UBL1 AOO87088.1     2
 Verru16b_00276 (CelZ)   Lacunisphaera limnophila IG16b AOS43233.1     2
 Lbys_2254   Leadbetterella byssophila DSM 17132 ADQ17931.1 E4RVN2   2
 clem_02275 (CelE)   Legionella clemsonensis CDC-D5610 ASQ45018.1     2
 LHA_2540   Legionella hackeliae ATCC35250 CEK11546.1     2
 SAMEA44548918_01078 (CelE)   Legionella lansingensis NCTC12830 SNV49872.1     2
 BBK82_45255   Lentzea guizhouensis DHS C013 ANZ42076.1     2
 LPB138_13260   Lutibacter sp. LPB0138 AOW21587.1     2
 LPB138_02890   Lutibacter sp. LPB0138 AOW19691.1     2
 Mahau_1113   Mahella australiensis 50-1 BON AEE96310.1     2
 BTR34_14425   Maribacter sp. T28 APQ18437.1     2
 CR152_18245   Massilia sp. B2 ATQ78963.1     2
 MROS_0753   Melioribacter roseus P3M-2 AFN73995.1     2
 Mag101_04990   Microbulbifer agarilyticus GP101 AQQ67068.1     2
 Mag101_04970   Microbulbifer agarilyticus GP101 AQQ67064.1     2
 A3224_06300   Microbulbifer thermotolerans DAU221 AMX02246.1     2
 Micau_2979   Micromonospora aurantiaca ATCC 27029 ADL46510.1 D9TBA5   2
 GA0070611_1781   Micromonospora auratinigra DSM 44815 SBT41892.1     2
 GA0070614_0094   Micromonospora coxensis DSM 45161 SCG34747.1     2
 GA0070610_3306   Micromonospora echinofusca DSM 43913 SCG17005.1     2
 GA0070618_5067   Micromonospora echinospora DSM 43816 SCF30627.1     2
 GA0070618_5917   Micromonospora echinospora DSM 43816 SCF37480.1     2
 GA0070621_1393   Micromonospora narathiwatensis DSM 45248 SBT42013.1     2
 GA0074696_3702   Micromonospora purpureochromogenes DSM 43821 SCF24260.1     2
 GA0070623_0785   Micromonospora rifamycinica DSM 44983 SCG41432.1     2
 GA0074704_3372   Micromonospora siamensis DSM 45097 SCG57304.1     2
 ML5_5408   Micromonospora sp. L5 ADU10877.1 D3C5S4   2
 CO540_05510   Micromonospora sp. WMMA2032 ATO13355.1     2
 GA0070619_3022   Micromonospora zamorensis DSM 45600 SCG53995.1     2
 A7982_04389   Minicystis rosea DSM 24000 APR79042.1     2
 SAMN05216490_0949   Mucilaginibacter mallensis MP1X4 SDS32391.1     2
 A0256_21225   Mucilaginibacter sp. PAMC 26640 AMR33776.1     2
 Niako_3530   Niastella koreensis GR20-10 AEV99831.1     2
 Niako_4179   Niastella koreensis GR20-10 AEW00453.1     2
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