Activities in Sub Family	Chitosanase (EC 3.2.1.132); Endo-β-1,4-glucanase (EC 3.2.1.4); Lichenase / endo-β-1,3-1,4-glucanase (EC 3.2.1.73); Endo-β-1,4-xylanase (EC 3.2.1.8);
Mechanism	Retaining e
Clan	GH-A
3D Structure Status	( β / α ) 8 barrel
Catalytic Nucleophile/Base	Glu (experimental)
Catalytic Proton Donor	Glu (experimental)
Note	Once known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resources	CAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)	MEGAZYME; NZYTech; PROZOMIX;
Statistics	GenBank accession (10); Uniprot accession (147); PDB accession (47); 3D entries (17); cryst (0)

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Bacteria
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D	Subf
MM221_10140		Salipaludibacillus sp. LMS25	UTR16833.1			2
RJT11_12325		Segatella copri H-2383	WOG02998.1			2
KUA49_000755		Segatella copri HDE06	WOZ84976.1			2
RJT13_07855		Segatella copri N-01	MDV3121554.1			2
BWZ22_15220		Seonamhaeicola sp. S2-3	APY12487.1			2
JYB87_11055		Shewanella avicenniae FJAT-51800	QSX32313.1			2
H9Q77_15745		Simiaoa sunii NSJ-8	QNM02472.1			2
cellulase (Cel5tm2)		Simiduia sp. TM-2	BAX64261.1			2
Solca_2501		Solitalea canadensis DSM 3403	AFD07536.1			2
O7608_20210		Solwaraspora sp. WMMA2056	WJK44020.1			2
O7553_12925		Solwaraspora sp. WMMA2059	WBB99719.1			2
O7610_07670		Solwaraspora sp. WMMA2065	WJK36223.1			2
O7543_04445		Solwaraspora sp. WMMA2080	WBC21731.1			2
O7623_03920		Solwaraspora sp. WMMD791	WFE28369.1			2
O7621_21435		Solwaraspora sp. WMMD937	WFE24641.1			2
LVJ94_07265 (fragment)		Sorangiineae bacterium MSr11367	WXB07031.1			2
LZC94_37245		Sorangiineae bacterium MSr11954	WXB13478.1			2
LZC95_06100		Sorangiineae bacterium MSr12523	WXA96409.1			2
LZC95_48590		Sorangiineae bacterium MSr12523	WXA94292.1			2
SOCE26_059880		Sorangium cellulosum So ce26	AUX44524.1			2
SOCE26_082810		Sorangium cellulosum So ce26	AUX46772.1			2
SOCE26_020140		Sorangium cellulosum So ce26	AUX40613.1			2
sce1468		Sorangium cellulosum So ce56	CAN91626.1			2
sce4910 (fragment)		Sorangium cellulosum So ce56	CAN95073.1			2
SOCE836_063270		Sorangium cellulosum So ce836	AUX34159.1			2
SOCE836_019920		Sorangium cellulosum So ce836	AUX29899.1			2
SOCE836_060420		Sorangium cellulosum So ce836	AUX33875.1			2
SOCEGT47_061770		Sorangium cellulosum So ceGT47	AUX25628.1			2
SOCEGT47_013300		Sorangium cellulosum So ceGT47	AUX20854.1			2
SCE1572_09655		Sorangium cellulosum So0157-2	AGP34753.1			2
SCE1572_28110		Sorangium cellulosum So0157-2	AGP37996.1			2
SCE1572_29465		Sorangium cellulosum So0157-2	AGP38247.1			2
NQZ70_01974		Sorangium sp. Soce836	WCQ89287.1			2
NQZ70_06219 (EglS)		Sorangium sp. Soce836	WCQ93470.1			2
NQZ70_05933		Sorangium sp. Soce836	WCQ93185.1			2
I6J33_12690		Sphingobacterium multivorum FDAARGOS_1203	QRQ63760.1			2
G6053_08115		Sphingobacterium sp. DR205	QIH37006.1			2
NCTC11429_04414 (EglS)		Sphingobacterium thalpophilum NCTC11429	VTR51732.1			2
PPZ50_01205		Sphingomonas hankookensis SZ.B2R2	WCP72213.1			2
G7077_04030		Sphingomonas piscis HDW15B	QIK79927.1			2
NMP03_15150		Sphingomonas qomolangmaensis S5-59	UUL82485.1			2
NMP03_15640 (fragment)		Sphingomonas qomolangmaensis S5-59	UUL82580.1			2
OKW76_13510		Sphingomonas sp. S1-29	UZK69031.1			2
ETR14_16160		Sphingosinicella sp. BN140058	QAY77883.1			2
STHERM_c14520		Spirochaeta thermophila DSM 6192	ADN02392.1			2
Spith_0779		Spirochaeta thermophila DSM 6578	AEJ61055.1			2
GJR95_13775		Spirosoma endbachense I-24	QHV96013.1			2
AWR27_01240		Spirosoma montaniterrae DY10	AQG78093.1			2
LQ777_07110		Spirosoma oryzicola RHs26	UHG92665.1			2
SAMN06298216_2068		Spirosomataceae bacterium TFI 002	SOE21610.1			2
Snas_1941		Stackebrandtia nassauensis DSM 44728	ADD41637.1			2
QU660_05010		Stomatobaculum sp. F0698	WLD85872.1			2
OUQ99_20925		Streptomonospora nanhaiensis 12A09	WAE71684.1			2
DMT42_01655		Streptomyces actuosus ATCC 25421	AWT41159.1			2
K1J60_43405		Streptomyces akebiae MG28	QYX82490.1			2
SMD44_00718		Streptomyces alboflavus MDJK44	ARX81320.1			2
QPM16_31185		Streptomyces anulatus K-31	WIY79648.1			2
OG771_32280		Streptomyces anulatus NBC_01421	WUY07998.1			2
KZO11_30660		Streptomyces anulatus VUW1	QYA97664.1			2
cellulase (Cel5A)		Streptomyces argenteolus M178	BAP19001.1			2
DVA86_04025		Streptomyces armeniacus ATCC 15676	AXK31937.1			2
CD934_32715		Streptomyces asterosporus DSM 41452	QDI72935.1			2
CD934_31065		Streptomyces asterosporus DSM 41452	QDI72641.1			2
G3H79_38970		Streptomyces aureoverticillatus HN6	QIB48175.1			2
chitosanase (OUC-Csngly)	3.2.1.4 3.2.1.132	Streptomyces bacillaris ATCC 15855	ULR56997.1			2
DN051_38390		Streptomyces cadmiisoli ZFG47	AWW41785.1			2
I6J39_29175		Streptomyces californicus FDAARGOS_1209	QRV32313.1			2
I6J40_05535		Streptomyces californicus FDAARGOS_1210	QRV58812.1			2
I6J41_29105		Streptomyces californicus FDAARGOS_1211	QRV45731.1			2
I6J42_04910		Streptomyces californicus FDAARGOS_1212	QRV38536.1			2
I6J43_28735		Streptomyces californicus FDAARGOS_1213	QRV52406.1			2
IHE65_43385		Streptomyces caniscabiei ID-03-3A	WEO29475.1			2
N0X72_09255		Streptomyces carpaticus SCPM-O-B-9993	UWM49195.1			2
CVT27_28035		Streptomyces cavourensis 1AS2a	ATY98913.1			2
NLU04_11815		Streptomyces cavourensis F2	UTR79100.1			2
OUQ49_28400		Streptomyces cavourensis NA4	WAE69374.1			2
DTW94_28310		Streptomyces cavourensis TJ430	AXI74767.1			2
NRO40_27670		Streptomyces changanensis HL-66	UUS34236.1			2
CP983_42705		Streptomyces chartreusis ATCC 14922	QEV72698.1			2
HUT05_46075		Streptomyces chartreusis NA02069	QKZ24069.1			2
CP977_31505		Streptomyces cinereoruber ATCC 19740	QEV36139.1			2
STRCI_008428		Streptomyces cinnabarinus DSM 40467	WAZ26790.1			2
CRV15_00710		Streptomyces clavuligerus ATCC 27064	QCS09154.1			2
LL058_27110		Streptomyces clavuligerus DSM 738	WDN55212.1			2
D1794_00710		Streptomyces clavuligerus F1D-5	AXU16371.1			2
I3J04_00610		Streptomyces clavuligerus F1D1	QPL61500.1			2
I3J07_00665		Streptomyces clavuligerus F1D10	QPL79642.1			2
I3J08_27435		Streptomyces clavuligerus F1D14	QPL90523.1			2
I3J09_00615		Streptomyces clavuligerus F1D18	QPL97605.1			2
I3J15_00590		Streptomyces clavuligerus F1D25	QPL91963.1			2
I3J16_00630		Streptomyces clavuligerus F1D26	QPM03618.1			2
I3J19_00610		Streptomyces clavuligerus F1D28	QPM21918.1			2
I3J17_27455		Streptomyces clavuligerus F1D31	QPM14391.1			2
I3J18_00645		Streptomyces clavuligerus F1D35	QPM15677.1			2
I3J06_00620		Streptomyces clavuligerus F1D7	QPL73609.1			2
BB341_00650		Streptomyces clavuligerus F613-1	ANW21741.1			2
SCO7637 or SC10F4.10c		Streptomyces coelicolor A3(2)	CAC16970.1 NP_631677.1			2
FQ762_39365		Streptomyces coelicolor A3(2) CFB_NBC_0001	QFI47356.1			2
JCM4020_76440		Streptomyces coelicolor JCM 4020	BDD77024.1			2
HCU77_38435		Streptomyces coelicolor M1154/pAMX4/pGP1416	QKN70841.1			2
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