Glycoside Hydrolase Family 5 / Subf 2

Activities in Sub Family	Chitosanase (EC 3.2.1.132); Endo-β-1,4-glucanase (EC 3.2.1.4); Lichenase / endo-β-1,3-1,4-glucanase (EC 3.2.1.73); Endo-β-1,4-xylanase (EC 3.2.1.8);
Mechanism	Retaining e
Clan	GH-A
3D Structure Status	( β / α ) ₈ barrel
Catalytic Nucleophile/Base	Glu (experimental)
Catalytic Proton Donor	Glu (experimental)
Note	Once known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resources	CAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)	MEGAZYME; NZYTech; PROZOMIX;
Statistics	GenBank accession (10); Uniprot accession (147); PDB accession (47); 3D entries (17); cryst (0)

Summary

Download GH5_2 (2298) Taxonomic display Structure (17) Characterized (129)

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Bacteria
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D	Subf
DI487_13690		Flavobacterium sediminis MEBiC07310	AWM14796.1			2
AABD74_21605		Flavobacterium soyae 55	WYZ19748.1			2
FK004_17195		Flavobacterium sp. AJ004	AWG26841.1			2
cellulase GH5		Flavobacterium sp. AUG42	QCD26741.1			2
HAV12_006320		Flavobacterium sp. CLA17	QSB28350.1			2
FLGSB24_42720		Flavobacterium sp. GSB-24	BDU27528.1			2
MKO97_13005		Flavobacterium sp. HJ-32-4	UMY65412.1			2
PQ462_18420		Flavobacterium sp. KACC 22758	WDF58689.1			2
SCB73_01295		Flavobacterium sp. KACC 22761	WPO79030.1			2
PQ463_12515		Flavobacterium sp. KACC 22763	WDF62447.1			2
B0M43_0023715		Flavobacterium sp. KBS0721	QDW23000.1			2
LPB248_16805		Flavobacterium sp. LPB0248 LPB248	QLC67898.1			2
HQN62_15445		Flavobacterium sp. M31R6	QKJ64469.1			2
IHE43_08235		Flavobacterium sp. MDT1-60	QOG04178.1			2
GIY83_00635		Flavobacterium sp. SLB02 SGTM	QGK72628.1			2
N4T20_21165		Flavobacterium sp. TR2	UWY28218.1			2
P0R33_17765		Flavobacterium sp. YJ01	WET01605.1			2
Q1W72_15415		Flavobacterium sp. ZE23DGlu08	WKL43724.1			2
LVD16_12735		Fulvivirga ligni W9P-11	UII24085.1			2
LVD15_25380		Fulvivirga maritima SW1-E11	UII26588.1			2
HX109_10185		Galbibacter sp. bg1	QLE01904.1			2
endo-β-1,4-glucanase (CelA;GsCelA)	3.2.1.4	Geobacillus sp. 70PC53	ACJ60856.1	C5H6X3	4XZB[A] 4XZW[A]	2
endoglucanase		Geobacillus stearothermophilus CICC-20649	ADO21451.1			2
NHM04_11975		Gilvimarinus sp. DA14	UTF59193.1			2
GCM10025732_34630		Glycomyces mayteni JCM 16217	BFF25498.1			2
GCM10025732_43900 (fragment)		Glycomyces mayteni JCM 16217	BFF26425.1			2
K3N28_10435		Glycomyces sp. TRM65418	QZD57487.1			2
SANA_14470		Gottschalkiaceae bacterium SANA	BES65008.1			2
Spica_2444		Gracilinema caldarium DSM 7334	AEJ20552.1			2
endoglucanase (Cel5A)		Gynuella sunshinyii YC6258	AII80614.1			2
YC6258_05810		Gynuella sunshinyii YC6258	AJQ97838.1			2
YC6258_02995		Gynuella sunshinyii YC6258	AJQ95033.1			2
endoglucanase (Cel5;HCH_03914)	3.2.1.4	Hahella chejuensis KCTC 2396	ABC30636.1	Q2SFD8		2
O5O45_07980		Hahella sp. HNIBRBA332	WLQ15851.1			2
ENC22_12810		Hahella sp. KA22	AZZ92039.1 QAY55410.1			2
endo-β-1,4-glucanase (Egl-1139) / III-3A (Cel5A)	3.2.1.4	Halalkalibacter akibai / KSM-1139	BAA00045.1 AAA22305.1 ACE14924.1	P06564		2
endoglucanase (EG-I-1)	3.2.1.4	Halalkalibacter akibai I-1	AFI25187.1	I1V1X7		2
SD1D_0239 (EglA)		Herbinix luporum SD1D	CUH91792.1			2
A4V00_16090		Hungateiclostridiaceae bacterium KB18	ANU55408.1			2
MUN79_25500		Hymenobacter cellulosilyticus 5116S-3	UOQ71911.1			2
MUN80_23325		Hymenobacter cellulosivorans 5116S-27	UOQ52664.1			2
DDQ68_08965		Hymenobacter nivis NBRC 111535	AWM35364.1			2
AXW84_14545		Hymenobacter sp. PAMC 26628	AMJ66514.1			2
Hsw_1037		Hymenobacter swuensis DY53	AHJ96632.1			2
MTX78_16550		Hymenobacter tibetensis KACC 21982	UOG73722.1			2
MTX78_16555		Hymenobacter tibetensis KACC 21982	UOG73723.1			2
MUN86_16370		Hymenobacter volaticus 5420S-77	UOQ65122.1			2
MUN86_25780		Hymenobacter volaticus 5420S-77	UOQ69129.1			2
BN2497_11653		Janthinobacterium sp. CG23_2	CUI24051.1			2
L0B70_12260		Kaistella sp. 97-N-M2	UJF29595.1			2
AOZ06_10145		Kibdelosporangium phytohabitans KLBMP1111	ALG07233.1			2
ORF		Kibdelosporangium sp. MJ126-NF4	CTQ95794.1			2
B7C62_30465		Kitasatospora albolonga YIM 101047	ARF76115.1			2
RGQ21_76160		Kitasatospora aureofaciens Tu117	BET52634.1			2
Kfla_1714		Kribbella flavida DSM 17836	ADB30809.1			2
OX958_27795		Kribbella sp. CA-293567	WBQ03764.1			2
V1226_12050		Lachnospiraceae bacterium JLR.KK009	WWR22054.1			2
cellulase, partial (Cel1) (fragment)		Lactiplantibacillus plantarum UBL1	AOO87088.1			2
extracellular cellulase, partial		Lactobacillus sp. UBRL-4	USS20256.1			2
Verru16b_00276 (CelZ)		Lacunisphaera limnophila IG16b	AOS43233.1			2
Lbys_2254		Leadbetterella byssophila DSM 17132	ADQ17931.1			2
PXX05_07575		Legionella cardiaca H63	WED41796.1			2
clem_02275 (CelE)		Legionella clemsonensis CDC-D5610	ASQ45018.1			2
LHA_2540		Legionella hackeliae ATCC35250	CEK11546.1			2
SAMEA44548918_01078 (CelE)		Legionella lansingensis NCTC12830	SNV49872.1			2
GH742_00170		Legionella sp. MW5194	QRN05007.1			2
FOLKNPGA_01884		Legionella sp. PC1000	QLZ69102.1			2
PQO03_21210		Lentisphaera profundi SAORIC-696	WDE98334.1			2
BBK82_45255		Lentzea guizhouensis DHS C013	ANZ42076.1			2
V1227_14225		Myxococcaceae DG1S-22	WVH83855.1			2
ND450_18045		Lentzea sp. HUAS12	USX55922.1			2
ND450_17080		Lentzea sp. HUAS12	USX55745.1			2
ATTO_11640		Leptogranulimonas caecicola TOC12	BDC91292.1			2
Mahau_1113		Mahella australiensis 50-1 BON	AEE96310.1			2
BTR34_14425		Maribacter hydrothermalis T28	APQ18437.1			2
MARGE09_P0761		Marinagarivorans cellulosilyticus GE09	BCD96561.1			2
MARGE09_P1526		Marinagarivorans cellulosilyticus GE09	BCD97325.1			2
MFFC18_09140 (CelZ)		Mariniblastus fucicola FC18	QEG21062.1			2
IV454_17950		Massilia antarctica P9640	QPI47491.1			2
CR152_18245		Massilia violaceinigra B2	ATQ78963.1			2
INH39_00275		Massilia violaceinigra sipir	UOD30238.1			2
MROS_0753		Melioribacter roseus P3M-2	AFN73995.1			2
MVE64_12310 (fragment)		Metabacillus endolithicus KCTC 33579	UPG65680.1			2
KDJ21_006395		Metabacillus litoralis NCTR108	UHA61285.1			2
LPC09_24455		Metabacillus sp. B2-18	UGB30803.1			2
LTA6_000610		Microbacterium sp. LTA6	WXH15255.1			2
Mag101_04990		Microbulbifer agarilyticus GP101	AQQ67068.1			2
Mag101_04970		Microbulbifer agarilyticus GP101	AQQ67064.1			2
BTJ40_12585		Microbulbifer sp. A4B17	AWF81594.1			2
endo-β-1,4-glucanase (MaCel5A) (Cel5A)	3.2.1.4	Microbulbifer sp. ALW1	QSR83793.1	A0A8A1G1R1		2
GL2_27000		Microbulbifer sp. GL-2	BBM02626.1			2
M8T91_04305		Microbulbifer sp. MI-G	WKD50656.1			2
FIU95_08260 (CelZ)		Microbulbifer sp. THAF38	QFT54545.1			2
A3224_06300		Microbulbifer thermotolerans DAU221	AMX02246.1			2
Q2E61_06420		Microbulbifer thermotolerans HB226069	WKT61824.1			2
Micau_2979		Micromonospora aurantiaca ATCC 27029	ADL46510.1			2
LF814_14950		Micromonospora aurantiaca G9	UFN97725.1			2
DVH21_22645		Micromonospora aurantiaca H14(2018)	AXH92495.1			2
GA0070611_1781		Micromonospora auratinigra DSM 44815	SBT41892.1			2
HZU44_16780		Micromonospora carbonacea africana	QLJ96590.1			2
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