Glycoside Hydrolase Family 5 / Subf 1

Activities in Familyendo-β-1,4-glucanase / cellulase (EC 3.2.1.4); endo-β-1,4-xylanase (EC 3.2.1.8); β-glucosidase (EC 3.2.1.21); β-mannosidase (EC 3.2.1.25); β-glucosylceramidase (EC 3.2.1.45); glucan β-1,3-glucosidase (EC 3.2.1.58); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); glucan endo-1,6-β-glucosidase (EC 3.2.1.75); mannan endo-β-1,4-mannosidase (EC 3.2.1.78); cellulose β-1,4-cellobiosidase (EC 3.2.1.91); steryl β-glucosidase (EC 3.2.1.104); endoglycoceramidase (EC 3.2.1.123); chitosanase (EC 3.2.1.132); β-primeverosidase (EC 3.2.1.149); xyloglucan-specific endo-β-1,4-glucanase (EC 3.2.1.151); endo-β-1,6-galactanase (EC 3.2.1.164); β-1,3-mannanase (EC 3.2.1.-); arabinoxylan-specific endo-β-1,4-xylanase (EC 3.2.1.-); mannan transglycosylase (EC 2.4.1.-); lichenase / endo-β-1,3-1,4-glucanase (EC 3.2.1.73); β-glycosidase (EC 3.2.1.-); endo-β-1,3-glucanase / laminarinase (EC 3.2.1.39); β-N-acetylhexosaminidase (EC 3.2.1.52); chitosanase (EC 3.2.1.132); β-D-galactofuranosidase (EC 3.2.1.146); β-galactosylceramidase (EC 3.2.1.46); ; β-rutinosidase /α-L-rhamnose-(1,6)-β-D-glucosidase (EC 3.2.1.-); α-L-arabinofuranosidase (EC 3.2.1.55)
Activities in Sub Family
Mechanism Retaining
ClanGH-A
3D Structure Status( β / α ) 8
Catalytic Nucleophile/BaseGlu (experimental)
Catalytic Proton DonorGlu (experimental)
NoteOnce known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)MEGAZYME; NZYTech; PROZOMIX;
Statistics GenBank accession (803); Uniprot accession (102); PDB accession (15); 3D entries (3); cryst (0)
All (763) Archaea (11) Bacteria (689) Eukaryota (50) Viruses (11) unclassified (2) Structure (3) Characterized (34)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 |
Bacteria
Protein Name EC#Organism GenBankUniprotPDB/3DSubf
 EGX37_11885   Clavibacter michiganensis subsp. nebraskensis 7580 QGV70268.1     1
 EGX36_11930   Clavibacter michiganensis subsp. nebraskensis HF4 QGV67468.1     1
 CMN_02390 (CeL)   Clavibacter michiganensis subsp. nebraskensis NCPPB 2581 CCE76326.1     1
 cellulase (CelA;pCS0061) 3.2.1.4 Clavibacter michiganensis subsp. sepedonicus ATCC33113 AAK16222.1
CAQ03244.1
B0RJ54
Q9AF65
  1
 CAC0912   Clostridium acetobutylicum ATCC 824 AAK78888.1
NP_347548.1
Q97KK6   1
 SMB_G0929 (CelB)   Clostridium acetobutylicum DSM 1731 AEI33087.1     1
 CEA_G0924   Clostridium acetobutylicum EA 2018 ADZ19963.1     1
 DK921_10960   Clostridium acetobutylicum LJ4 AWV80607.1     1
 endoglucanase (Eng5A)   Clostridium cellulovorans BAV13030.1     1
 endoglucanase (Eng5E)   Clostridium cellulovorans BAV13035.1     1
 Clocel_0619   Clostridium cellulovorans 743B ADL50390.1 D9SRM3   1
 Clocel_0983   Clostridium cellulovorans 743B ADL50749.1 D9STD3   1
 Cspa_c18680 (Celb2)   Clostridium saccharoperbutylacetonicum N1-4(HMT) AGF55638.1     1
 Clo1100_1829   Clostridium sp. BNL1100 AEY66039.1     1
 Clo1100_2788   Clostridium sp. BNL1100 AEY66946.1     1
 B5808_07170   Cnuibacter physcomitrellae XA(T) ARJ05010.1     1
 HH215_30930   Cohnella sp. MFER-1 QJD87157.1     1
 Cwoe_0159   Conexibacter woesei DSM 14684 ADB48595.1 D3F517   1
 PA27867_1437   Cryobacterium arcticum PAMC 27867 ANP72394.1     1
 Cyan10605_2560   Cyanobacterium aponinum PCC 10605 AFZ54640.1     1
 CYFUS_005260   Cystobacter fuscus DSM 52655 ATB39812.1     1
 EUS_18130   [Eubacterium] siraeum 70/3 CBK96886.1 D4JUW7   1
 ES1_23790   [Eubacterium] siraeum V10Sc8a CBL35203.1 D4MN70   1
 C1H87_01040   Flavivirga eckloniae ECD14 AUP77378.1     1
 cellulase GH5   Flavobacterium sp. AUG42 QCD26736.1     1
 SAMN04488543_1663   Friedmanniella luteola DSM 21741 SDS40102.1     1
 SAMN04488544_1121 (fragment)   Friedmanniella sagamiharensis DSM 21743 SDU86229.1     1
 GM3709_2278   Geminocystis sp. NIES-3709 BAQ65513.1     1
 Glo7428_3546   Gloeocapsa sp. PCC 7428 AFZ32015.1     1
 A9404_07265   Halothiobacillus sp. LS2 ANJ67208.1     1
 HL652_05860   Herbiconiux sp. SALV-R1 QJU53200.1     1
 HL652_11565   Herbiconiux sp. SALV-R1 QJU56062.1     1
 SD1D_0238 (Celb2)   Herbinix luporum SD1D CUH91791.1     1
 SD1D_0139 (CelB)   Herbinix luporum SD1D CUH91693.1     1
 cellulase (H4;Haur_2820)   Herpetosiphon aurantiacus DSM 785 ABX05458.1 A9B2K7   1
 DVJ78_13720   Humibacter sp. BT305 AXH36339.1     1
 endoglucanase Cel5   Hungateiclostridium cellulolyticum WP_010249057.1     1
 Clocl_1981   Hungateiclostridium clariflavum DSM 19732 AEV68583.1     1
 Clocl_3932   Hungateiclostridium clariflavum DSM 19732 AEV70372.1     1
 Clocl_1122   Hungateiclostridium clariflavum DSM 19732 AEV67797.1     1
 HVS_03380   Hungateiclostridium saccincola GGR1 AUG56624.1     1
 HVS_07025   Hungateiclostridium saccincola GGR1 AUG57322.1     1
 AD2_02839   Hungateiclostridium thermocellum AD2 ALX09817.1     1
 AD2_01719   Hungateiclostridium thermocellum AD2 ALX08709.1     1
 AD2_00416   Hungateiclostridium thermocellum AD2 ALX07424.1     1
 AD2_01837   Hungateiclostridium thermocellum AD2 ALX08827.1     1
 Cthe_0405 (CelL)   Hungateiclostridium thermocellum ATCC 27405 ABN51643.1 A3DCG4   1
 endo-β-1,4-glucanase B (CelB;EgB;Cthe_0536) (Cel5A) 3.2.1.4 Hungateiclostridium thermocellum ATCC 27405 ABN51772.1 P04956   1
 Cthe_2147 (CelO) (Cel5F)   Hungateiclostridium thermocellum ATCC 27405 ABN53350.1 A3DHC1   1
 endo-β-1,4-glucanase G (CelG;Cthe_2872) (Cel5D) 3.2.1.4 Hungateiclostridium thermocellum ATCC 27405 ABN54070.1 Q05332   1
 Clo1313_1701   Hungateiclostridium thermocellum DSM 1313 ADU74758.1     1
 Clo1313_1816   Hungateiclostridium thermocellum DSM 1313 ADU74872.1     1
 Clo1313_2805   Hungateiclostridium thermocellum DSM 1313 ADU75785.1     1
 Clo1313_0413   Hungateiclostridium thermocellum DSM 1313 ADU73502.1     1
 LQRI_1836   Hungateiclostridium thermocellum DSM 2360 LQRI ANV76577.1     1
 LQRI_1720   Hungateiclostridium thermocellum DSM 2360 LQRI ANV76461.1     1
 LQRI_2850   Hungateiclostridium thermocellum DSM 2360 LQRI ANV77591.1     1
 LQRI_0416   Hungateiclostridium thermocellum DSM 2360 LQRI ANV75163.1     1
 endo-1,4-β-glucanase (CelL)   Hungateiclostridium thermocellum F7 CAK18680.1 Q1JR84   1
 cellobiohydrolase (CelO) (Cel5F) 3.2.1.91 Hungateiclostridium thermocellum F7 CAB76938.1 Q9L3J2   1
 endo-β-1,4-glucanase B (CelB;EgB) (Cel5A) 3.2.1.4 Hungateiclostridium thermocellum NCIB 10682 CAA27266.1 P04956   1
 endo-β-1,4-glucanase G (CelG) (Cel5D) 3.2.1.4 Hungateiclostridium thermocellum NCIB 10682 CAA49187.1 Q05332   1
 BN2497_13425   Janthinobacterium sp. CG23_2 CUI24937.1     1
 Jden_0734   Jonesia denitrificans DSM 20603 ACV08398.1 C7R1T0   1
 CEP80_01495   Jonesia denitrificans FDAARGOS_301 ASE07953.1     1
 NCTC10816_00755 (EngxcA)   Jonesia denitrificans NCTC10816 SQH20376.1     1
 endoglucanase, partial (Cel5A)   Jonesia quinghaiensis Y2 API85532.1     1
 KSE_69000   Kitasatospora setae KM-6054 BAJ32658.1 E4N3C4   1
 endo-β-1,4-glucanase 5A (CpCel5A;Cphy_1163) (Cel5A) 3.2.1.4 Lachnoclostridium phytofermentans ISDg ABX41541.1 A9KN42   1
 Lxx22675 (CelA)   Leifsonia xyli subsp. xyli str. CTCB07 AAT89935.1 Q6ACF8   1
 LBDG_00070   Leptolyngbya boryana dg5 BAS60474.1     1
 LBWT_70 (fragment)   Leptolyngbya boryana IAM M-101 BAS54126.1     1
 NIES2135_06290   Leptolyngbya boryana NIES-2135 BAY53817.1     1
 LEP3755_40970   Leptolyngbya sp. NIES-3755 BAU13557.1     1
 O77CONTIG1_02914   Leptolyngbya sp. O-77 BAU43092.1     1
 Mahau_1116   Mahella australiensis 50-1 BON AEE96313.1     1
 AZF01_06880   Martelella sp. AD-3 AMM84112.1     1
 CR152_25160   Massilia violaceinigra B2 ATQ77421.1     1
 C6568_02270   Melaminivora sp. SC2-9 AVO48212.1     1
 MEBOL_000684   Melittangium boletus DSM 14713 ATB27246.1     1
 DK389_16305   Methylobacterium durans 17SD2-17 AWN41780.1     1
 GNH96_02885   Methylococcus sp. IM1 QJD29018.1     1
 BWL13_02425 (fragment)   Microbacterium oleivorans A9 AZS44828.1     1
 SAMN04489810_3464   Microbacterium pygmaeum DSM 23142 SDH61822.1     1
 BJP65_02895   Microbacterium sp. BH-3-3-3 AOX44867.1     1
 SAMN05216488_1924   Microbacterium sp. LKL04 SCY46623.1     1
 CEP17_14150   Microbacterium sp. PM5 AXA97461.1     1
 MTES_0141   Microbacterium testaceum StLB037 BAJ73105.1     1
 Mic7113_4470   Microcoleus sp. PCC 7113 AFZ20162.1     1
 Micau_1546   Micromonospora aurantiaca ATCC 27029 ADL45104.1 D9SYR9   1
 DVH21_15565   Micromonospora aurantiaca H14(2018) AXH91240.1     1
 GA0070611_6160   Micromonospora auratinigra DSM 44815 SBT53509.1     1
 GA0070612_5967   Micromonospora chokoriensis DSM 45160 SCF29189.1     1
 GA0070607_1055   Micromonospora coriariae DSM 44875 SCE74509.1     1
 GA0070614_4776   Micromonospora coxensis DSM 45161 SCG71042.1     1
 GA0070614_0384   Micromonospora coxensis DSM 45161 SCG37348.1     1
 GA0070610_4046   Micromonospora echinofusca DSM 43913 SCG17725.1     1
 GA0070618_0392   Micromonospora echinospora DSM 43816 SCE71858.1     1
 GA0070621_0342   Micromonospora narathiwatensis DSM 45248 SBT38231.1     1
 GA0070623_2127   Micromonospora rifamycinica DSM 44983 SCG53214.1     1
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