Activities in Sub Family	Endo-β-1,4-glucanase (EC 3.2.1.4); Lichenase / endo-β-1,3-1,4-glucanase (EC 3.2.1.73); Exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74);
Mechanism	Retaining e
Clan	GH-A
3D Structure Status	( β / α ) 8 barrel
Catalytic Nucleophile/Base	Glu (experimental)
Catalytic Proton Donor	Glu (experimental)
Note	Once known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resources	CAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)	MEGAZYME; NZYTech; PROZOMIX;
Statistics	GenBank accession (59); Uniprot accession (38); PDB accession (16); 3D entries (4); cryst (0)

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Bacteria
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D	Subf
T3H97_18725		halophilic eubacterium NRCC 41227 LX16	WVD75094.1			1
C0638_14995		Paenibacillus sp. lzh-N1	AUO07745.1			1
EGM68_18755		Paenibacillus sp. M-152	AZH30655.1			1
EI981_10975		Paenibacillus sp. MBLB1234	AZS14931.1			1
LOS79_21345		Paenibacillus sp. MMS20-IR301	WNS41556.1			1
KAI36_03904		Paenibacillus sp. S02	QYK68751.1			1
KAI37_03839		Paenibacillus sp. S25	QYK63506.1			1
HPL003_26960		Paenibacillus terrae HPL-003	AET62104.1			1
KDC22_32015		Paenibacillus tritici PH55-1	QUL54817.1			1
Acav_3053		Paracidovorax avenae ATCC 19860	ADX46955.1			1
R1Z03_09665		Paracidovorax avenae BLS4	WOI47447.1			1
C8240_13915		Paracidovorax cattleyae CAT98_1	AVS74938.1			1
Aave_2102		Paracidovorax citrulli AAC00-1	ABM32683.1			1
LKW27_08130		Paracidovorax citrulli HPP21-3-3B	UEG47817.1			1
QRO11_08340		Paracidovorax citrulli KACC 17001	WIY36323.1			1
QRO08_08440		Paracidovorax citrulli KACC 17005	WIY50579.1			1
QRO12_16430		Paracidovorax citrulli KACC 17913	WIY42533.1			1
QRO09_04650		Paracidovorax citrulli KACC 18782	WIY31019.1			1
QRO10_04590		Paracidovorax citrulli KACC 18784	WIY40230.1			1
CQB05_04045		Paracidovorax citrulli KACC17005	ATG93317.1			1
APS58_0225 (EngxcA)		Paracidovorax citrulli M6	QCX09195.1			1
FRC75_04700		Paracidovorax citrulli NWB SC074	UMT82738.1			1
FRC90_13280		Paracidovorax citrulli NWB SC107	UMT88938.1			1
FRC97_15795		Paracidovorax citrulli NWB SC196	UMT96342.1			1
endoglucanase, partial		Paracidovorax citrulli xjgb-wy6	AKT09126.1			1
SAMN04489860_0019		Paraoerskovia marina DSM 22126	SDR77692.1			1
EQG49_04655		Periweissella cryptocerci 26KH-42	QBO35805.1			1
C7M71_023240		Peterkaempfera bronchialis DSM 106435	AXI79881.1			1
Pflav_088110		Phytohabitans flavus NBRC 107702	BCB82401.1			1
Psuf_051100		Phytohabitans suffuscus NBRC 105367	BCB87797.1			1
PBC10988_21160		Planctomycetales bacterium 10988	QGJ70419.1			1
Pan216_00890		Planctomycetes bacterium Pan216	QDU59261.1			1
FXF36_01895		Pseudobutyrivibrio xylanivorans MA3014	QFJ53706.1			1
HZ992_12960		Rhizobacter sp. AJA081-3	QTN21119.1			1
DLJ82_7530		Rhizobium leguminosarum ATCC 14479	AXA43775.1			1
LZK79_26630		Rhizobium leguminosarum Tp73_4	UIK20460.1			1
RlegTA1_35580		Rhizobium ruizarguesonis TA1	UFW99237.1			1
RHOA_1246		Rhodovastum atsumiense G2-11	CAH2599851.1			1
Riv7116_4185		Rivularia sp. PCC 7116	AFY56617.1			1
Riv7116_0722		Rivularia sp. PCC 7116	AFY53310.1			1
N4261_13030		Roseateles amylovorans BIM B-1768	UXH80740.1			1
RD2015_747		Roseateles depolymerans KCTC 42856	ALV05243.1			1
RD2015_749		Roseateles depolymerans KCTC 42856	ALV05245.1			1
OU995_01610		Roseateles sp. SL47	WAC73472.1			1
OU995_01600		Roseateles sp. SL47	WAC73470.1			1
GFK88_29955		Roseibium aggregatum ZYF703	UES54007.1			1
Rmf_34940		Roseomonas fluvialis S08	BDG73565.1			1
RGI145_02340		Roseomonas gilardii U14-5	APT56123.1			1
ADP8_00830		Roseomonas mucosa AD1	QDD98961.1			1
RMP42_00830		Roseomonas mucosa AD3	UZO91154.1			1
FOB66_03555		Roseomonas mucosa FDAARGOS_658	QET92012.1			1
WDZ11_12625		Roseomonas mucosa GlSt084	WXH21879.1			1
RMHFA_00830 (fragment)		Roseomonas mucosa HV1	UZO96007.1			1
HVIM_00830		Roseomonas mucosa HV3	QDD93857.1			1
RADP37_00830		Roseomonas mucosa P37	AWV21756.1			1
HVPorG_00830		Roseomonas mucosa PG	QDJ08674.1			1
NF552_12320		Roseomonas mucosa TT14	USQ73582.1			1
LPC08_02160		Roseomonas sp. OT10	UFN49473.1			1
CMCase (P66;Ccel_2337)	3.2.1.4	Ruminiclostridium cellulolyticum H10	ACL76673.1	B8I5D7		1
cellulase N (CelN; Ccel_0740) (Cel5N)	3.2.1.4	Ruminiclostridium cellulolyticum H10	ACL75118.1 AAG45162.1	B8I7W1 Q9EYQ0		1
EHE19_014380		Ruminiclostridium herbifermentans MA18	QNU66059.1			1
EHE19_001790		Ruminiclostridium herbifermentans MA18	QNU67298.1			1
Cel5B (Cel5B)		Ruminiclostridium josui	BAI52933.1			1
P0092_15785		Ruminiclostridium papyrosolvens DSM 2782	WES33211.1			1
P0092_10495		Ruminiclostridium papyrosolvens DSM 2782	WES36372.1			1
Rumal_1049		Ruminococcus albus 7 = DSM 20455	ADU21575.1			1
Rumal_2017		Ruminococcus albus 7 = DSM 20455	ADU22510.1			1
Rumal_2777		Ruminococcus albus 7 = DSM 20455	ADU23246.1			1
cellulase (Cel5G)		Ruminococcus albus 8	AAT48117.1			1
lichenase (Ra2830;CUS_5609)	3.2.1.73	Ruminococcus albus 8	EGC04285.1	E9S8W4		1
RBI_I00497		Ruminococcus bicirculans (ex Wegman et al. 2014) 80/3	CCO04221.1			1
endo-β-1,4-glucanase (GH5B;RUM_06330) (Cel5B)	3.2.1.4	Ruminococcus champanellensis 18P13 = JCM 17042	CBL16847.1	D4LB52		1
GCM10017745_63620		Saccharothrix mutabilis subsp. capreolus JCM 4248	BFE52935.1			1
IOD16_15240		Saccharothrix sp. 6-C	QQQ79626.1			1
L3Q67_29620		Saccharothrix sp. AJ9571	UJW29380.1			1
EKG83_24680		Saccharothrix syringae NRRL B-16468	QFZ20180.1			1
Strop_2684		Salinispora tropica CNB-440	ABP55128.1			1
SAMD00079811_23510		Scytonema sp. HK-05 NIES-2130	BAY44749.1			1
SAMD00079811_39570		Scytonema sp. HK-05 NIES-2130	BAY46348.1			1
NIES4073_72930		Scytonema sp. NIES-4073	BAZ26387.1			1
NIES4073_56600		Scytonema sp. NIES-4073	BAZ24761.1			1
LHU95_20980		Sediminicoccus rosea KRV36	UPY36670.1			1
LHU95_19005		Sediminicoccus rosea KRV36	UPY36285.1			1
R9Z33_11570		Sediminicoccus rosea R-30	WPB87492.1			1
R9Z33_01995		Sediminicoccus rosea R-30	WPB85657.1			1
SVI_0875		Shewanella violacea DSS12	BAJ00846.1			1
M5M_14915		Simiduia agarivorans SA1 = DSM 21679	AFV00118.1 AFV00107.2			1
JL100_021190		Skermanella mucosa KEMB 2255-438	UEM19585.1			1
JL101_019905		Skermanella rosea KEMB 2255-458	UEM02247.1			1
IGS68_06410		Skermanella sp. TT6	QQP90854.1			1
O7608_15845		Solwaraspora sp. WMMA2056	WJK43977.1			1
O7553_16620		Solwaraspora sp. WMMA2059	WBC00341.1			1
O7610_11120		Solwaraspora sp. WMMA2065	WJK36842.1			1
O7543_00755		Solwaraspora sp. WMMA2080	WBC21072.1			1
O7623_09665		Solwaraspora sp. WMMD791	WFE29428.1			1
O7621_25000		Solwaraspora sp. WMMD937	WFE21081.1			1
LZC95_04605		Sorangiineae bacterium MSr12523	WXA96118.1			1
STHERM_c20620 (StX;StCBM64A)		Spirochaeta thermophila DSM 6192	ADN02996.1			1
Spith_2109		Spirochaeta thermophila DSM 6578	AEJ62365.1			1
GXW83_22715		Streptacidiphilus sp. PB12-B1b	QMU78089.1			1
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