Activities in Family	β-glucosidase (EC 3.2.1.21); β-galactosidase (EC 3.2.1.23); β-mannosidase (EC 3.2.1.25); β-glucuronidase (EC 3.2.1.31); β-xylosidase (EC 3.2.1.37); β-D-fucosidase (EC 3.2.1.38); phlorizin hydrolase (EC 3.2.1.62); exo-β-1,4-glucanase (EC 3.2.1.74); 6-phospho-β-galactosidase (EC 3.2.1.85); 6-phospho-β-glucosidase (EC 3.2.1.86); strictosidine β-glucosidase (EC 3.2.1.105); lactase (EC 3.2.1.108); amygdalin β-glucosidase (EC 3.2.1.117); prunasin β-glucosidase (EC 3.2.1.118); vicianin hydrolase (EC 3.2.1.119); raucaffricine β-glucosidase (EC 3.2.1.125); thioglucosidase (EC 3.2.1.147); β-primeverosidase (EC 3.2.1.149); isoflavonoid 7-O-β-apiosyl-β-glucosidase (EC 3.2.1.161); ABA-specific β-glucosidase (EC 3.2.1.175); DIMBOA β-glucosidase (EC 3.2.1.182); hydroxyisourate hydrolase (EC 3.-.-.-); β-rutinosidase /α-L-rhamnose-(1,6)-β-D-glucosidase (EC 3.2.1.-); protodioscin 26-O-β-D-glucosidase (EC 3.2.1.186); α-L-arabinopyranosidase (EC 3.2.1.-); isoflavone 7-O-glucosyl β-glucosidase (EC 3.2.1.-)
Mechanism	Retaining
Clan	GH-A
3D Structure Status	( β / α ) 8 barrel
Catalytic Nucleophile/Base	Glu (experimental)
Catalytic Proton Donor	Glu (experimental); absent in plant myrosinases
External resources	CAZypedia; HOMSTRAD; PRINTS; PROSITE;
Commercial Enzyme Provider(s)	MEGAZYME; NZYTech; PROZOMIX;
Statistics	GenBank accession (58998); Uniprot accession (452); PDB accession (300); 3D entries (81); cryst (0)

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Archaea
Protein Name	EC#	Reference	Organism	GenBank	Uniprot	PDB/3D
multifunctional β-glycosidase (β-Gly;Asβ-Gly;ASAC_1390)	3.2.1.21 3.2.1.23	pubmed pubmed	Acidilobus saccharovorans 345-15	ADL19795.1	D9PZ08	4HA3[A] 4HA4[A]
Cmaq_0408	3.2.1.21	pubmed	Caldivirga maquilingensis IC-167	ABW01253.1	A8MBR0
Cmaq_0652	3.2.1.21	pubmed	Caldivirga maquilingensis IC-167	ABW01492.1	A8MCI6
β-galactosidase (CelB)	3.2.1.23	pubmed	Pyrococcus furiosus B232	AHW49177.1	Q51723
β-mannosidase (BglB;PF0356)	3.2.1.25	pubmed	Pyrococcus furiosus DSM 3638	NP_578085.1 AAL80480.1	Q8U3U9
β-glucosidase / β-rutinosidase (Cel1A;CelB;BGLPf;PfBgl1;PFTG;PF0073;Pf-β-gly)	3.2.1.21 3.2.1.-	pubmed pubmed	Pyrococcus furiosus DSM 3638	AAL80197.1 AAC25555.1 ABC06396.1 NP_577802.1	E7FHY4 Q51723	3APG[A,B,C,D] 3WDP[P,Q,R,S] 3WQ8[A,B,C,D,E,F,G,H,I,J,K,L]
β-mannosidase / β-galactosidase (BmnA;PF1208)	3.2.1.23 3.2.1.25	pubmed pubmed	Pyrococcus furiosus DSM 3638	AAL81332.1 AAC44387.1 AAM56789.1 ABC06395.1 CAH68690.1 CAT81449.1 NP_578937.1	E7FHC0 Q51733
β-glucosidase (BglA;PF0442) (Cel1B)	3.2.1.21	pubmed	Pyrococcus furiosus DSM 3638	AAL80566.1 AAG28457.1 NP_578171.1	E7FI48 Q9HHB3
alkyl β-glucosidase (membrane-bound) (BGPh;BglA;PH0366)	3.2.1.21	pubmed	Pyrococcus horikoshii OT3	BAA29440.1 ABC06393.1 NP_142340.1	O58104	1VFF[A]
β-mannosidase (BglB;PH0501)	3.2.1.25	pubmed	Pyrococcus horikoshii OT3	BAA29589.1 ABC06394.1 NP_142473.1	O58237
β-galactosidase	3.2.1.23	pubmed	Pyrococcus woesei	AAB97862.1	O52629
β-glycosidase (Bgly;SSG)	3.2.1.21	pubmed	Saccharolobus shibatae	AAA79030.1	P50388
β-glycosidase / β-galactosidase	3.2.1.21 3.2.1.23	doi doi	Saccharolobus solfataricus DSM1616	CAA34074.1	P14288
β-glycosidase S (LacS; Ss-β-gly; SSO3019)	3.2.1.21 3.2.1.23	pubmed pubmed	Saccharolobus solfataricus P2	AAA72843.1 AAE29896.1 AAK43121.1 ABC06397.1 ACW34039.1 CAC24042.1 CAJ18790.1 NP_344331.1	P22498	1GOW[A,B] 1UWI[A,B,C,D] 1UWQ[A,B] 1UWR[A,B] 1UWS[A,B] 1UWT[A,B] 1UWU[A,B] 2CEQ[A,B] 2CER[A,B] 4EAM[A,B] 4EAN[A,B] 5I3D[A,B,C,D] 5IXE[A,B]
β-glycosidase (BgaS;Saci_1849)	3.2.1.21	pubmed	Sulfolobus acidocaldarius DSM 639	AAY81155.1 YP_256448.1	P14288
β-glycosidase (TK1827)	3.2.1.21 3.2.1.23	pubmed pubmed	Thermococcus kodakarensis KOD1	BAD86016.1 CAT70991.1	Q5JDV2
β-glycosidase (Tkbgly;TK1761)	3.2.1.21 3.2.1.23	pubmed pubmed	Thermococcus kodakarensis KOD1	BAA78713.1 BAD85950.1 CAT70557.1	Q5JDT7 Q9YGB8
β-glucosidase (BglT)	3.2.1.21	pubmed	Thermococcus sp. AL662	CAA94187.1	O08324
β-glucosidase (TsBGL; B6U94_07015)	3.2.1.21	pubmed	Thermofilum sp. ex4484_79	OYT30282.1		7F1N[A,B]
TVG0691226	3.2.1.21 3.2.1.23	pubmed pubmed	Thermoplasma volcanium GSS1	BAB59827.1 NP_111204.1	Q97AX4
β-glucosidase (Tα-β-gly;Gly)	3.2.1.21	pubmed	Thermosphaera aggregans M11TL	AAD43138.1 AAM56782.1	Q9YGA8	1QVB[A,B]
β-glucosidase (Vul_Bgl1A)	3.2.1.21	pubmed	uncultured archaeon	QGN01269.1
β-glucosidase (Bgl1)	3.2.1.108 3.2.1.21 3.2.1.38	pubmed pubmed pubmed	uncultured archaeon	CDG15325.1	W8W3B8
Bacteria
Protein Name	EC#	Reference	Organism	GenBank	Uniprot	PDB/3D
exo-β-glucosidase 1A (BglA; β-G;Cthe_0212) (Glc1A)	3.2.1.21	pubmed	Acetivibrio thermocellus ATCC 27405	CAA42814.1 AAQ79758.1 ABN51453.1	P26208	5OGZ[A,B]
Acel_0133	3.2.1.21	pubmed	Acidothermus cellulolyticus 11B	ABK51908.1	A0LR48
β-glucosidase (Abg)	3.2.1.21	pubmed	Agrobacterium sp. ATCC21400	AAA22085.1 AAQ79763.1	P12614
β-glucosidase (AacGH1)	3.2.1.21	pubmed	Alicyclobacillus acidiphilus	WP_067621817.1		6ZIV[AAA,BBB,CCC,DDD,EEE,FFF,GGG,HHH]
β-glycosidase (GlyB;Aaci_1895)	3.2.1.21 3.2.1.23	pubmed pubmed	Alicyclobacillus acidocaldarius subsp. acidocaldarius DSM 446	ACV58907.1 AAZ81839.1	C8WXU5 Q0R5R7
β-glucosidase	3.2.1.21 3.2.1.38 3.2.1.-	pubmed pubmed pubmed	Alicyclobacillus herbarius		A0A8I3B065	6YN7[A,B,C,D]
Aflv_2360	3.2.1.21	pubmed	Anoxybacillus flavithermus WK1	ACJ34717.1	B7GM33
β-glucosidase (BglZ)	3.2.1.21	pubmed	Bacillus amyloliquefaciens ABBD	AFS64714.1	J9ZZW8
BLi04199 / BL01946	3.2.1.21	pubmed	Bacillus licheniformis DSM 13 = ATCC 14580	AAU43012.1 AAU25633.1 CBG21160.1	Q65D52
6-P-β-glucosidase (BglH; BlBglH; BLi04214; BL00212)	3.2.1.86	pubmed	Bacillus licheniformis DSM 13 = ATCC 14580	AAU43027.1 AAU25648.1	Q65D37	6WGD[A,B,C]
BLi00387 or BL01718 (YckE)	3.2.1.38	pubmed	Bacillus licheniformis DSM 13 = ATCC 14580	AAU39345.1 AAU21991.1 CBG21156.1	Q65NL9	7M1R[A,B,C,D]
β-glucosidase (Bgl1A; BsBgl1A)	3.2.1.21	pubmed	Bacillus sp. CGMCC 1.16541	QMX85751.1 WP_110112460.1	A0A7G5WHW4
β-glucosidase A	3.2.1.21	pubmed	Bacillus sp. GL1	BAA36160.1 CBG21242.1	Q9ZNN7
aryl β-glucosidase (BglC)	3.2.1.21	pubmed	Bacillus sp. SJ-10	AHC94275.1	V9VGK4
6-P-β-glucosidase (YdhP;BSU05840)	3.2.1.86	pubmed	Bacillus subtilis subsp. subtilis str. 168	BAA19708.1 AIY91871.1 AQR80536.1 AQR84749.1 CAB12403.1 CBG21028.1 NP_388465.1	A0A6M3Z8A5 O05508
6-P-β-glucosidase (BglA;BSU40110)	3.2.1.86	pubmed	Bacillus subtilis subsp. subtilis str. 168	AAA22660.1 AIY95329.1 AQR84054.1 AQR88269.1 BAA11270.1 CAB16048.1 CBG21250.1 NP_391891.1	A0A6M4JR50 P42973
6-P-β-glucosidase 1 (BglH;N17D;BSU39260)	3.2.1.86	pubmed	Bacillus subtilis subsp. subtilis str. 168	CAB15962.2 AIY95585.1 AQR83968.1 AQR88183.1 CAA84287.1 CBG21247.1 NP_391805.1	A0A6M3ZHY9 P40740
6-P-β-glucosidase (YckE;BglC;BSU03410)	3.2.1.86	pubmed	Bacillus subtilis subsp. subtilis str. 168	CAB12135.1 AIY91623.1 AQR80279.1 AQR84492.1 BAA06429.1 BAA08975.1 CBG21248.1 NP_388223.1	A0A6M3ZAF5 P42403
β-glucosidase / β-xylosidase (BglB;P11-6B)	3.2.1.21 3.2.1.37	pubmed pubmed	bacterium enrichment culture clone P11-6B	ADT62000.1	E9KYU3
β-glucosidase / β-D-fucosidase	3.2.1.21 3.2.1.38	pubmed pubmed	Bifidobacterium breve 203 / clb	BAA19881.1 CBG21199.1	P94248
β-glucosidase / cellodextrinase (CldC; Bbr_0109)	3.2.1.21	pubmed	Bifidobacterium breve UCC2003	ACT78497.1 ABE94821.1	F1LL52
β-glucosidase (Bgl13;Calkr_2207)	3.2.1.21	URL	Caldicellulosiruptor acetigenus I77R1B	ADQ41669.1	E4S6B1
β-glucosidase / cellodextrinase (CbBgl1A;CbCdx1A;Athe_0458) (Bgl1A=Cdx1A)	3.2.1.21	doi	Caldicellulosiruptor bescii DSM 6725	ACM59590.1	B9MNR1
β-glucosidase A (BglA)	3.2.1.21	pubmed	Caldicellulosiruptor saccharolyticus	CAA31087.1 AAQ79759.1 CBG21257.1	P10482
β-glucosidase (BglA;Csac_1089)	3.2.1.21	pubmed	Caldicellulosiruptor saccharolyticus DSM 8903	ABP66702.1	A4XIG7
β-galactosidase	3.2.1.23	doi	Caldicellulosiruptor sp. enrichment culture clone YTY-70	AFQ98284.1	J7KFN3
β-glucosidase (BlgA)	3.2.1.21	pubmed	Caldicellulosiruptor sp. F32	AFO70070.1	I7DLX2
CCNA_02220	3.2.1.21	pubmed	Caulobacter vibrioides NA1000	ACL95685.1	A0A0H3C9V1 B8GXX8
cellobiase (Cba3)	3.2.1.21	pubmed	Cellulomonas biazotea ATCC 486	AEM45802.1	G1C1T5
β-glucosidase (CfBgl1;Celf_2783)	3.2.1.21	pubmed	Cellulomonas fimi ATCC 484	AEE46907.1	F4H7E6
Cfla_1084	3.2.1.21	pubmed	Cellulomonas flavigena DSM 20109	ADG73989.1	D5ULE7
Rsph17025_1146	3.2.1.21	pubmed	Cereibacter sphaeroides ATCC 17025	ABP70047.1	A4WRN3
β-glucosidase A (BglA;BG)	3.2.1.21	pubmed	Clostridium cellulovorans	AAQ00997.1 CBG21209.1	Q53EH2	3AHX[A,B,C,D]
6-P-β-glucosidase (AbgA)	3.2.1.86	pubmed	Clostridium longisporum	AAC05714.1	Q46130
ESA_02547	3.2.1.86	pubmed	Cronobacter sakazakii ATCC BAA-894	ABU77792.1	A7MJ23
Deide_00550	3.2.1.21	pubmed	Deinococcus deserti VCD115	ACO44852.1	C1CXP6
β-glucosidase A (BglA;DP2448)	3.2.1.21	pubmed	Desulfotalea psychrophila LSv54	CAG37177.1 YP_066184.1	Q6AKE8
β-glucosidase (DtGH;DICTH_1689)	3.2.1.21	pubmed	Dictyoglomus thermophilum H-6-12	ACI19973.1	B5YAN1
β-glucosidase (Dtur?Glu; Dtur_0462)	3.2.1.21	pubmed	Dictyoglomus turgidum DSM 6724	ACK41762.1	B8E1X9
β-galactosidase (Dtur_1799)	3.2.1.21	pubmed	Dictyoglomus turgidum DSM 6724	ACK43071.1	B8E0Q1
6-P-β-glucosidase (AscB;ECs3572)	3.2.1.86	pubmed	Escherichia coli O157:H7 str. Sakai	BAB36995.1 NP_311599.1	Q8X841
6-P-β-glucosidase (BglB;b3721)	3.2.1.86	pubmed	Escherichia coli str. K-12 substr. MG1655	AAC76744.1 AAA23511.1 AAA62072.1 AAA83838.1 AAQ79757.1 AIZ93475.1 ALI38586.1 AMC96613.1 AUG18433.1 AVI53891.1 AYG21461.1 NP_418177.1	P11988
6-P-β-glucosidase 2 (AscB;b2716)	3.2.1.86	pubmed	Escherichia coli str. K-12 substr. MG1655	AAA69226.1 AAA16430.1 AAC75758.1 AIZ93701.1 ALI42228.1 AMC98974.1 AUG19188.1 AVI56782.1 AYG21513.1 NP_417196.1	P24240
6-P-β-glucosidase A (BglA;b2901)	3.2.1.86	pubmed	Escherichia coli str. K-12 substr. MG1655	AAC75939.1 AAA83082.1 AUG17619.1 AVI53071.1 AYG18126.1 CBG20988.1 NP_417377.1	Q46829	2XHY[A,B,C,D]
β-glucosidase (BglA;EaBglA;EaBgl1A;Eab7_0297)	3.2.1.21	pubmed	Exiguobacterium antarcticum B7 Eab7	AFS69459.1	K0A8J9	5DT5[A,B,C,D,E,F,G,H] 5DT7[A,B,C,D]
EAT1b_2183	3.2.1.21	doi	Exiguobacterium sp. AT1b	ACQ71106.1	C4L1S4	6WIU[A,B,C,D,E,F,G,H,I,J,K,L,M,N,O,P]
β-glucosidase (BglA)	3.2.1.21	doi	Exiguobacterium sp. DAU5	ADK47980.1	E0X9H4
β-glucosidase (FiBgl1A) (Bgl1A)	3.2.1.21	pubmed	Fervidobacterium islandicum	CCA60742.1	G8YZD7
Fnod_1558	3.2.1.21	pubmed	Fervidobacterium nodosum Rt17-B1	ABS61401.1	A7HNB8
β-glucosidase (BglA)	3.2.1.21	pubmed	Fervidobacterium sp. YNP	AAN60220.1	Q8GKQ8
6-P-β-glucosidase (PbgA)	3.2.1.86	pubmed	Fusobacterium mortiferum	AAB49339.1	P94793
β-galactosidase (GKGal1A;GK2337) (Gal1A)	3.2.1.23	doi	Geobacillus kaustophilus HTA426	BAD76622.1 CBG21168.1	Q5KXG4
6-phospho-β-glycosidase (GK3214)	3.2.1.85 3.2.1.86	pubmed pubmed	Geobacillus kaustophilus HTA426	BAD77499.1 CBG21169.1	Q5KUY7
6-phospho-β-glycosidase (GK1856)	3.2.1.85 3.2.1.86	pubmed pubmed	Geobacillus kaustophilus HTA426	BAD76141.1 CBG21167.1	Q5KYU5
6-P-β-galactosidase (Gan1D)	3.2.1.85 3.2.1.86	pubmed pubmed	Geobacillus stearothermophilus T-1	AHL67640.1	W8QF82	4ZE4[A,B] 4ZE5[A,B,C,D] 4ZEH[A,B,C,D] 4ZEN[A,B] 4ZEP[A,B] 4ZFM[A,B,C,D] 5OK7[A,B,C,D] 5OKA[A,B,C,D] 5OKB[A,B,C,D] 5OKE[A,B,C,D] 5OKG[A,B,C,D] 5OKH[A,B] 5OKJ[A,B] 5OKK[A,B] 5OKQ[A,B] 5OKR[A,B] 5OKS[A,B] 8B80[A,B] 8B81[A,B]
HCH_06907	3.2.1.21	pubmed	Hahella chejuensis KCTC 2396	ABC33525.1	Q2S749
β-glucosidase (BglA;BH1923)	3.2.1.21	pubmed	Halalkalibacterium halodurans C-125	BAB05642.1 CBG20998.1 NP_242789.1	Q9KBK3
β-galactosidase (BgalH)	3.2.1.23	pubmed	Halomonas sp. S62	AFA35120.1	H6WX44
β-glucosidase A (BglA;HoBGLA;Hore_15280)	3.2.1.21	pubmed	Halothermothrix orenii H 168	ACL70277.1	B8CYA8	3TA9[A,B] 4PTV[A,B] 4PTW[A,B] 4PTX[A,B]
β-glucosidase (BglD5; JMA_34740)	3.2.1.21		Jeotgalibacillus malaysiensis D5	AJD92791.1	A0A0B5ARU7
6-P-β-glucosidase (BglB)	3.2.1.86	pubmed	Klebsiella aerogenes AN292	AAM82757.1	Q8KP24
6-P-β-glucosidase / phospho-cellobiase (CasB)	3.2.1.86	pubmed	Klebsiella oxytoca P2	AAB51564.1 CBG21258.1	Q48409
[ginsenoside] β-glucosidase (KfGH01W; Kfla_5268)	3.2.1.21	pubmed	Kribbella flavida DSM 17836	ADB34282.1		6M6L[A,B] 6M6M[A,B]
6-P-β-galactosidase	3.2.1.85	pubmed	Lacticaseibacillus casei	AAD15134.1 AAA25238.1 AAQ79754.1 CAB02557.1 CBG21260.1	P14696
aryl 6-P-β-glucosidase (BglH;BglT)	3.2.1.86	pubmed	Lactiplantibacillus plantarum B21	CAA75906.1	O86291
6-P-β-galactosidase	3.2.1.85	doi	Lactobacillus acidophilus	BAA07122.1 CBG21259.1	P50977
6-P-β-glucosidase (Pbg1;LGAS_0185)	3.2.1.86	pubmed	Lactobacillus gasseri ATCC 33323 = JCM 1131	ABJ59596.1	A0A830PMB8 Q046M6
6-P-β-galactosidase (LacG1;LGAS_0344)	3.2.1.85	pubmed	Lactobacillus gasseri ATCC 33323 = JCM 1131	ABJ59750.1	A0A805ZFN4 Q045X2
6-P-β-galactosidase (LacG2;LGAS_0502)	3.2.1.85	pubmed	Lactobacillus gasseri ATCC 33323 = JCM 1131	ABJ59900.1	A0A805ZG68 Q045H2
6-P-β-glucosidase (Pbg2;LGAS_1668)	3.2.1.86	pubmed	Lactobacillus gasseri ATCC 33323 = JCM 1131	ABJ60960.1	A0A805ZJ83 Q041B2
6-P-β-galactosidase 1 (Pbg1)	3.2.1.86	pubmed	Lactobacillus gasseri JCM1031	BAA20086.1 BAD95552.1 CBG21210.1	O06532 Q588H0
6-P-β-galactosidase II (Pbg2)	3.2.1.86	pubmed	Lactobacillus gasseri JCM1031	BAA25004.1 CBG21197.1	O50658
llmg_1046	3.2.1.86	pubmed	Lactococcus cremoris subsp. cremoris MG1363	CAL97639.1	A2RK29
6-P-β-galactosidase (LacG)	3.2.1.85	pubmed	Lactococcus lactis Z268	AAA25183.1 AAA25173.1 AAA26949.1 AAQ79752.1 CBG21261.1	P11546	1PBG[A,B] 2PBG[A] 3PBG[A,B] 4PBG[A,B]
amb0219	3.2.1.21	pubmed	Magnetospirillum magneticum AMB-1	BAE49023.1	Q2WAV2
β-glucosidase (BglM1;Mmwyl1_4295) (cold-active)	3.2.1.21	doi	Marinomonas sp. MWYL1	ABR73190.1	A6W3B1
β-glucosidase (Mrub_0288)	3.2.1.21	pubmed	Meiothermus ruber DSM 1279	ADD27066.1 AGK03521.1	D3PLV5
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