Activities in Family	[retaining] anthocyanin 5-O-glucosyltransferase / 5-O-transglucosidase (EC 2.4.1.-); [retaining] anthocyanin 7-O-glucosyltransferase / 7-O-transglucosidase (EC 2.4.1.-); [retaining] β-transglucosidase (EC 2.4.1.-); α-L-arabinopyranosidase (EC 3.2.1.-); β-1,2-glucosidase (EC 3.2.1.-); β-rutinosidase /α-L-rhamnose-(1,6)-β-D-glucosidase (EC 3.2.1.-); Isoflavone 7-O-glucosyl β-glucosidase (EC 3.2.1.-); 3-a-(S)-strictosidine β-glucosidase (EC 3.2.1.105); Lactase (EC 3.2.1.108); Amygdalin β-glucosidase (EC 3.2.1.117); Prunasin β-glucosidase (EC 3.2.1.118); [retaining] vicianin hydrolase (EC 3.2.1.119); Raucaffricine β-glucosidase (EC 3.2.1.125); Myrosinase (EC 3.2.1.147); β-primeverosidase (EC 3.2.1.149); β-acuminosidase / β-apiosyl β-glucosidase / furcatin hydrolase (EC 3.2.1.161); ABA-specific β-glucosidase (EC 3.2.1.175); Exo-α-sialidase (EC 3.2.1.18); DIMBOA β-glucosidase (EC 3.2.1.182); Protodioscin 26-O-β-D-glucosidase (EC 3.2.1.186); 0-26-avenacosidase (EC 3.2.1.188); Oleuropein β-glucosidase (EC 3.2.1.206); β-glucosidase (EC 3.2.1.21); β-galactosidase (EC 3.2.1.23); β-mannosidase (EC 3.2.1.25); β-glucuronidase (EC 3.2.1.31); Xylan β-1,4-xylosidase (EC 3.2.1.37); β-D-fucosidase (EC 3.2.1.38); β-glucosylceramidase (EC 3.2.1.45); β-galactosylceramidase (EC 3.2.1.46); Exo-β-1,3-glucanase (EC 3.2.1.58); Exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); 6-P-β-galactosidase (EC 3.2.1.85); 6-P-β-glucosidase (EC 3.2.1.86);
Mechanism	Retaining e
Clan	GH-A
3D Structure Status	( β / α ) 8 barrel
Catalytic Nucleophile/Base	Glu (experimental)
Catalytic Proton Donor	Glu (experimental); absent in plant myrosinases
External resources	CAZypedia; HOMSTRAD; PRINTS; PROSITE;
Commercial Enzyme Provider(s)	MEGAZYME; NZYTech; PROZOMIX;
Statistics	GenBank accession (66130); Uniprot accession (452); PDB accession (319); 3D entries (88); cryst (0)

Archaea
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D Carbohydrate Ligands Resolution (Å)
multifunctional β-glycosidase (β-Gly; Asβ-Gly; ASAC_1390)	3.2.1.21 3.2.1.23	Acidilobus saccharovorans 345-15	ADL19795.1	D9PZ08	4HA3[A] 1.46 4HA4[A] 1.37
β-glucosidase / β-rutinosidase (Cel1A;CelB;BGLPf;PfBgl1;PFTG;PF0073;Pf-β-gly)	3.2.1.21 3.2.1.-	Pyrococcus furiosus DSM 3638	AAL80197.1 AAC25555.1 ABC06396.1 NP_577802.1	E7FHY4 Q51723	3APG[A,B,C,D] 2.35 3WDP[P,Q,R,S] 1.70 3WQ8[A,B,C,D,E,F,G,H,I,J, 2.81
alkyl β-glucosidase (membrane-bound) (BGPh;BglA;PH0366)	3.2.1.21	Pyrococcus horikoshii OT3	BAA29440.1 ABC06393.1 NP_142340.1	O58104	1VFF[A] 2.50
β-glycosidase S (LacS; Ss-β-gly; SSO3019)	3.2.1.21 3.2.1.23	Saccharolobus solfataricus P2	AAA72843.1 AAE29896.1 AAK43121.1 ABC06397.1 ACW34039.1 CAC24042.1 CAJ18790.1 NP_344331.1	P22498	1GOW[A,B] 2.60 1UWI[A,B,C,D] 2.55 1UWQ[A,B] 2.02 1UWR[A,B] α-D-Galp2F-(1-4)-GLU 2.14 1UWS[A,B] α-D-Glcp2F-(1-4)-GLU 1.95 1UWT[A,B] <carb_like_GTL> 1.95 1UWU[A,B] <carb_like_GOX> 1.95 2CEQ[A,B] <carb_like_GIM> 2.14 2CER[A,B] <carb_like_PGI> 2.29 4EAM[A,B] 1.70 4EAN[A,B] 1.75 5I3D[A,B,C,D] 2.16 5IXE[A,B] 1.75 7UZ1[A,B] 1.58 7UZ2[A,B] 1.83
β-glucosidase (TsBGL; B6U94_07015)	3.2.1.21	Thermofilum sp. ex4484_79	OYT30282.1		7F1N[A,B] 2.14
β-glucosidase (Tα-β-gly;Gly)	3.2.1.21	Thermosphaera aggregans M11TL	AAD43138.1 AAM56782.1	Q9YGA8	1QVB[A,B] 2.40
Bacteria
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D Carbohydrate Ligands Resolution (Å)
exo-β-glucosidase 1A (BglA; β-G; HtBgl; Cthe_0212) (Glc1A)	3.2.1.21	Acetivibrio thermocellus ATCC 27405	CAA42814.1 AAQ79758.1 ABN51453.1	P26208	5OGZ[A,B] 1.60 8IVY[A,B] β-D-Glcp 1.95
SghA		Agrobacterium tumefaciens A6	APY21444.1		6RJK[A,B] 1.92 6RJM[A,B] α-D-Glcp 2.11 6RJO[A,B] β-D-Glcp-(1-1)-<non_carb> 1.80 6RK2[A,B] α-L-Altp-(1-1)-<non_carb> 2.09
β-glucosidase (AacGH1)	3.2.1.21	Alicyclobacillus acidiphilus	WP_067621817.1		6ZIV[AAA,BBB,CCC,DDD,EEE,
β-glucosidase (AheGH1)	3.2.1.21 3.2.1.38 3.2.1.-	Alicyclobacillus herbarius		A0A8I3B065	6YN7[A,B,C,D] 1.98
6-P-β-glucosidase (BglH; BlBglH; BLi04214; BL00212)	3.2.1.86	Bacillus licheniformis DSM 13 = ATCC 14580	AAU43027.1 AAU25648.1	Q65D37	6WGD[A,B,C] 2.20
BLi00387 or BL01718 (YckE; BlBglC)	3.2.1.38	Bacillus licheniformis DSM 13 = ATCC 14580	AAU39345.1 AAU21991.1 CBG21156.1	Q65NL9	7M1R[A,B,C,D] 1.98
β-4,6-pyryvylated-galactosidase (ORF1119)		Bacillus sp. (in: firmicutes) HMA207	BBF98480.1		5YHS[A,B] 2.50 5YIF[A,B] 2.45
CQJ30_16875		Caldibacillus thermoamylovorans SSBM	AWI14273.1		8HT0[A] 2.00
β-glucosidase A (BglA;BG)	3.2.1.21	Clostridium cellulovorans	AAQ00997.1 CBG21209.1	Q53EH2	3AHX[A,B,C,D] 1.90
6-P-β-glucosidase A (BglA;b2901)	3.2.1.86	Escherichia coli str. K-12 substr. MG1655	AAC75939.1 AAA83082.1 AUG17619.1 AVI53071.1 AYG18126.1 CBG20988.1 NP_417377.1	Q46829	2XHY[A,B,C,D] 2.30
β-glucosidase (BglA;EaBglA;EaBgl1A;Eab7_0297)	3.2.1.21	Exiguobacterium antarcticum B7 Eab7	AFS69459.1	K0A8J9	5DT5[A,B,C,D,E,F,G,H] 2.24 5DT7[A,B,C,D] 2.15
EAT1b_2183	3.2.1.21	Exiguobacterium sp. AT1b	ACQ71106.1	C4L1S4	6WIU[A,B,C,D,E,F,G,H,I,J, 3.51
Ferpe_1843		Fervidobacterium pennivorans DSM 9078	AFG35894.1		6KDC[A] 1.85
bifunctional 6-P-β-galactosidase / 6-P-β-glucosidase (Gan1D)	3.2.1.85 3.2.1.86	Geobacillus stearothermophilus T-1	AHL67640.1	W8QF82	4ZE4[A,B] 1.92 4ZE5[A,B,C,D] 1.48 4ZEH[A,B,C,D] 1.33 4ZEN[A,B] P-(0-6)-β-D-Galp 1.93 4ZEP[A,B] P-(0-6)-β-D-Glcp 2.21 4ZFM[A,B,C,D] P-(0-6)-β-D-Glcp P-(0-6)-β-D-Glcp-(1-4)-β-D-Glcp 1.40 5OK7[A,B,C,D] 1.34 5OKA[A,B,C,D] 1.25 5OKB[A,B,C,D] 1.33 5OKE[A,B,C,D] P-(0-6)-β-D-Glcp P-(0-6)-β-D-Glcp-(1-4)-β-D-Glcp 1.31 5OKG[A,B,C,D] P-(0-6)-β-D-Glcp P-(0-6)-β-D-Glcp-(1-4)-β-D-Glcp 1.25 5OKH[A,B] 1.98 5OKJ[A,B] 1.76 5OKK[A,B] P-(0-6)-β-D-Galp 2.02 5OKQ[A,B] P-(0-6)-β-D-Galp 1.85 5OKR[A,B] P-(0-6)-β-D-Glcp 2.15 5OKS[A,B] P-(0-6)-β-D-Glcp 1.96 8B80[A,B] P-(0-6)-β-D-Galp 1.78 8B81[A,B] P-(0-6)-β-D-Glcp-(1-4)-β-D-Glcp 1.59
β-glucosidase A (BglA;HoBGLA;Hore_15280)	3.2.1.21	Halothermothrix orenii H 168	ACL70277.1	B8CYA8	3TA9[A,B] 3.00 4PTV[A,B] β-D-Glcp-(1-4)-β-D-Glcp4S 1.85 4PTW[A,B] α-D-Glcp2F-(1-4)-GLU 2.00 4PTX[A,B] β-D-Glcp 1.80
[ginsenoside] β-glucosidase (KfGH01W; Kfla_5268)	3.2.1.21	Kribbella flavida DSM 17836	ADB34282.1		6M6L[A,B] β-D-Xylp 1.50 6M6M[A,B] 2.07
β-galactosidase (Bgl; lp_3629)	3.2.1.23	Lactiplantibacillus plantarum WCFS1	CAD65701.1 AAS68346.1 CBG21089.1 CCC80585.1 NP_786823.1		5NAQ[A,B,C,D,E,F] 2.48 5NAV[A,B,C,D,E,F] 2.30
β-galactosidase (Pbg1; LpPbg1; lp_0440)	3.2.1.23	Lactiplantibacillus plantarum WCFS1	CAD63073.1 CBG21079.1 CCC77945.1 NP_784234.1		3QOM[A] β-D-Glcp 1.50 4GZE[A,B,C,D,E,F] 2.31
6-P-β-galactosidase (LacG)	3.2.1.85	Lactococcus lactis Z268	AAA25183.1 AAA25173.1 AAA26949.1 AAQ79752.1 CBG21261.1	P11546	1PBG[A,B] 2.30 2PBG[A] 2.50 3PBG[A,B] 2.70 4PBG[A,B] P-(0-6)-β-D-Galp 2.50
β-glycosidase (TY87_18135)	3.2.1.21 3.2.1.23	Marinomonas sp. BSi20584	PJE53865.1		8PUO[A,B]
β-glycosidase (BGL167;BglG167c)	3.2.1.21 3.2.1.-	Microbacterium sp. Gsoil167	AGA60135.1	L0ELG0	4R27[A,B] 2.03
β-glucosidase (Bgl;BglU)	3.2.1.21	Micrococcus antarcticus	ACM66669.1 AMB49877.1 AMB49878.1 AMB49879.1	A0A0Y0GAW6 A0A0Y0KCV2 A0A0Y0M4F4 B9V8P5	3W53[A] 2.20
β-glucosidase (BglA)	3.2.1.21	Niallia circulans subsp. alkalophilus	AAA22266.1 AAQ79761.1 CBG21249.1	Q03506	1QOX[A] 2.7
β-glucosidase A (BglA)	3.2.1.21	Paenibacillus polymyxa	AAA22263.1 AAQ79760.1 CBG21252.1	P22073	1BGA[A,B,C,D] 2.40 1BGG[A,B,C,D] 2.30 1E4I[A] α-D-Glcp2F-(1-4)-GLU β-D-Glcp2F-(1-1)-<non_carb> 2.00 1TR1[A,B,C,D] 2.20 1UYQ[A] α-D-Glcp2F-(1-4)-GLU β-D-Glcp2F-(1-1)-<non_carb> 2.20 6QWI[A,B] 2.85 6R4K[A,B] 2.13
β-glucosidase B (BglB)	3.2.1.21	Paenibacillus polymyxa	AAA22264.1 AAQ79762.1 CBG21256.1	P22505	2JIE[A] α-D-Glcp2F-(1-4)-GLU 2.30 2O9P[A] 2.10 2O9R[A] β-D-Glcp-(1-4)-β-D-Glcp4S 2.30 2O9T[A] β-D-Glcp 2.15 2Z1S[A] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 2.46
6-P-β-glucosidase (CelA;Bgl;SmBgl;SMU.1601;SMU_1601)	3.2.1.86	Streptococcus mutans UA159	AAN59243.1 CBG21068.1 NP_721937.1	Q8DT00	3PN8[A,B] 1.69 4DDE[A,B] P-(0-6)-β-D-Glcp 1.45 4F66[A,B] P-(0-6)-β-D-Glcp 1.48 4F79[A] P-(0-6)-β-D-Glcp-(1-1)-<non_carb> 2.54 4GPN[A,B] P-(0-6)-β-D-Glcp-(1-6)-β-D-Glcp 2.29
6-P-β-glucosidase (BglA-2; SP0578)	3.2.1.86	Streptococcus pneumoniae TIGR4	AAK74732.1 CBG21011.1 NP_345092.1	A0A0H2UP35 Q97S37	4IPL[A,B] 2.00 4IPN[B,E] P-(0-6)-α-L-Idop-(1-4)-β-D-Glcp4S 2.41
6-phospho-β-glucosidase (SPy_1599;SPy1599)	3.2.1.86	Streptococcus pyogenes M1 GAS SF370	AAK34377.1 NP_269656.1	Q99YP9	4B3K[A,B,C,D,E,F] 2.60 4B3L[A,B,C,D,E,F] 2.51 5FOO[A,B,C,D,E,F] 2.10
β-glucosidase (Bgl3)	3.2.1.21	Streptomyces sp. QM-B814	CAA82733.1 CBG21211.1	Q59976	1GNX[A,B] α-D-Glcp-(1-2)-β-D-Fruf 1.68 1GON[A,B] 2.2
β-glucosidase (TsaBgl; Tsac_2208)	3.2.1.21	Thermoanaerobacterium saccharolyticum JW/SL-YS485	AFK87212.1		7E5J[A] 1.71 8WFT[A,B,C,D] 1.90 8WFU[A,B,C,D] 2.10 8WFV[A] 1.50 8WFW[A] 1.61 8XPC[A] 1.55 8XPD[A] 1.70 8XPE[A] 1.95
β-glucosidase A (BglA; Tmari_1862)	3.2.1.21	Thermotoga maritima MSB8	CAA52276.1 AEP25088.1 AGL50786.1 AHD18256.1 AKE29603.1 CBG21253.1	G4FGN9 G4XU74 Q08638 R4NU76	1OD0[A,B] 2.11 1OIF[A,B] <carb_like_IFM> 2.12 1OIM[A,B] <carb_like_NOJ> 2.15 1OIN[A,B] α-D-Glcp2F-(1-4)-GLU 2.15 1UZ1[A,B] <carb_like_IFL> 2.0 1W3J[A,B] <carb_like_OXZ> 2.0 2CBU[A,B] <carb_like_CTS> 1.85 2CBV[A,B] <carb_like_CGB> 1.95 2CES[A,B] <carb_like_GIM> 2.15 2CET[A,B] <carb_like_PGI> 1.97 2J75[A,B] <carb_like_NOY> 1.85 2J77[A,B] <carb_like_NOJ> 2.1 2J78[A,B] <carb_like_GOX> 1.65 2J79[A,B] <carb_like_GTL> 1.94 2J7B[A,B] <carb_like_NTZ> 1.87 2J7C[A,B] <carb_like_IDE> 2.09 2J7D[A,B] <carb_like_GI1> 2.24 2J7E[A,B] 2.19 2J7F[A,B] <carb_like_GI3> 2.28 2J7G[A,B] <carb_like_GI4> 1.91 2J7H[A,B] <carb_like_AZF> 1.95 2JAL[A,B] <carb_like_YLL> 1.9 2VRJ[A,B] 1.90 2WBG[A,B,C,D] <carb_like_LGS> 1.85 2WC3[A,B,C,D] <carb_like_AM3> 2.00 2WC4[A,B,C,D] <carb_like_AMF> 1.70 5N6S[A,B,C,D] 2.10 5N6T[A,B] 2.10 5OSS[A,B] 1.70
β-glucosidase A / 1A (BglA;GghA;TnBgl1A;CTN_0782) (Bgl1A)	3.2.1.21	Thermotoga neapolitana DSM 4359	AAB95492.2 ACM22958.1 CAB10165.1 CBG21254.1	B9K7M5 O33843 P0C946	5IDI[A,B] 1.90
β-glycosidase (Gly)	3.2.1.21	Thermus nonproteolyticus HG102	AAF36392.1	Q9L794	1NP2[A,B] 2.40
β-glycosidase / β-glucosidase (TTHB087)	3.2.1.21	Thermus thermophilus HB8	AAN05439.1 BAD71883.1 CBG21166.1	Q53W75 Q8GEB3	1UG6[A] 0.99 4BCE[A,B,C] 2.00
β-glycosidase (BglT;B-gly)	3.2.1.21 3.2.1.23	Thermus thermophilus TH125	AAD32630.2	Q9RA61	3ZJK[A,B,C] 2.20
β-glucosidase (Bgl6;BGL6)	3.2.1.21	uncultured bacterium	AKH41028.1	A0A0F7KKB7	5GNX[A,B] β-D-Glcp 1.80 5GNY[A,B,C,D] β-D-Glcp 2.11 5GNZ[A,B,C,D,H,I,J,K] β-D-Glcp 2.20
β-glucosidase Bg10		uncultured bacterium	APG42678.1		7BBS[A,B] 2.30
exo-β-1,3-glucanase	3.2.1.58	uncultured bacterium	APM84071.1		8J3M[A,B,C,D] 2.00 8J5L[A,B,C,D] 2.10 8J5M[A,B,C,D] 1.62
β-glucosidase (Bgl1317)	3.2.1.21	uncultured bacterium	QCG75868.1	A0A4D6T7S3 A0A5H1ZR35	6IER[A] 2.25
β-glucosidase (Bgl15)	3.2.1.21	uncultured bacterium	QED88370.1	A0A5B9BHU3	6JFP[A,B] β-D-Glcp 2.70
β-glucosidase / β-glycosidase (BglA) (Bgl1A)	3.2.1.21	uncultured bacterium	ABI18350.1	Q0GMU3	3CMJ[A] 1.60 3FIY[A] 1.40 3FIZ[A] β-D-Glcp 2.00 3FJ0[A] β-D-Glcp 1.15 4HZ6[A] 1.40 4HZ7[A] β-D-Glcp 2.00 4HZ8[A] β-D-Glcp 1.14
Eukaryota
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D Carbohydrate Ligands Resolution (Å)
β-glucosidase (AkuBGL; ApBGL; AkLab)		Aplysia kurodai	BAX08664.1		8IN1[A,B] 2.70
β-glucosidase (AkuBGL; ApBGL; AkLab)	3.2.1.21	Aplysia kurodai	BAX08664.1		8IN1[A,B] 2.70
myrosinase (TGG1;Bglu38;At5g26000)	3.2.1.147	Arabidopsis thaliana	AAC18869.1 AAD40143.1 AAK74039.1 AAL25596.1 AAL91284.1 AAN60236.1 AED93511.1 AED93512.1 BAH20253.1 CAA55786.1 CAH40799.1 CAH40800.1 CAH40801.1 CAH40802.1 CAH40803.1 CAH40804.1 CAH40805.1 CAH40806.1 CAH40807.1 CAH40808.1 CAH40809.1 CAH40810.1 CAH40811.1 CAH40812.1 CAH40813.1 CAH40814.1 CAH40815.1 CAH40816.1 CAH40817.1 CAH40818.1 CAH40819.1 CAH40820.1 CAH40821.1 CAH40822.1 CAH40823.1 CAH40824.1 CAH40825.1 CAH40826.1 NP_197972.1 NP_851077.1	A0A178U7L7 P37702	7Z1I[A,B,C,D] 3.09
β-glucosidase (Bglu42; At5g36890)	3.2.1.21	Arabidopsis thaliana	NP_198505.2 AAQ89633.1 AED94122.1 AED94123.1 BAB11630.1 BAD43523.1 CBG20994.1 NP_198505.1	A0A654G6E3 Q9FIW4	7F3A[A] 1.70
myrosinase (BMY1)	3.2.1.147	Brevicoryne brassicae	AAL25999.1 CBG21232.1	Q95X01	1WCG[A,B] 1.10
β-primeverosidase (Bgl;PD)	3.2.1.149	Camellia sinensis	BAC78656.1 AAR76593.1 AAR76595.1 AAS83105.1 AAS83105.2 ACH11663.1 ADV40931.1 ADV40931.2 AHM10169.1 ANQ46489.1 CAK97604.1 CAK97604.2 CBG21219.1	A0A1B1FKH7 E7EF69 Q14QP8 Q6PW57 Q7X9A9 W8SAR2	3WQ4[A,B] 1.90 3WQ5[A,B] β-D-Xylp-(1-6)-β-D-Glcp1N-(1-1)-<non_carb> 1.80 3WQ6[A,B] β-D-Xylp-(1-6)-β-D-Glcp1N-(1-1)-<non_carb> 1.80
β-galactosidase/β-glucosidase (BglA)	3.2.1.21 3.2.1.23	Hamamotoa singularis ATCC 24193	BAD95570.1	Q564N5	6M4E[A] 2.10 6M4F[A,C,E,G] 2.20 6M55[A,D] β-D-Galp-(1-4)-β-D-Galp-(1-4)-α-D-Glcp 3.00 7L74[A,B] 2.25
klotho (KL)		Homo sapiens	BAA23382.1 AAQ41828.1 AAQ41829.1 ACP57343.1 ACP57344.1 BAA24940.1 BAA24941.1 BAG53456.1 CAH71888.1 CAR95139.1 CAR95140.1		5W21[A] 3.00 7YSH[A] α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3 2.74 7YSU[A] α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GalpNSO36SO3-(1-4)-α-L-AltpA2SO3-(1-4)-α-D-GalpNSO36SO3-(1-4)-α-L-AltpA2SO3-(1-4)-α-D-GalpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3 3.20 7YSW[C] α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-AltpA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-AltpA2SO3-(1-4)-α-D-GalpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-AltpA2SO3 3.03
klotho (KL)		Homo sapiens	BAA23382.1 AAQ41828.1 AAQ41829.1 ACP57343.1 ACP57344.1 BAA24940.1 BAA24941.1 BAG53456.1 CAC94767.1 CAH71888.1 CAR95139.1 CAR95140.1		5W21[A] 3.00 7YSH[A] α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3 2.74 7YSU[A] α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GalpNSO36SO3-(1-4)-α-L-AltpA2SO3-(1-4)-α-D-GalpNSO36SO3-(1-4)-α-L-AltpA2SO3-(1-4)-α-D-GalpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3 3.20 7YSW[C] α-D-GlcpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-AltpA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-AltpA2SO3-(1-4)-α-D-GalpNSO36SO3-(1-4)-α-L-IdopA2SO3-(1-4)-α-D-GlcpNSO36SO3-(1-4)-α-L-AltpA2SO3 3.03
neutral β-glycosylceramidase / β-glucosidase (Gba3;CbgCBgl1;GluC;KLrP) (cytosolic)	3.2.1.21 3.2.1.45	Homo sapiens	CAC08178.1 AAG39217.1 AAH29362.1 AAI01830.1 AAI09378.1 AAL37305.1 BAB18741.1 BAD96683.1 BAF85435.1 BAG36006.1 BAH12772.1 NP_066024.1	A8K9N1 B7Z536 Q9H227	2E9L[A] β-D-Glcp 1.60 2E9M[A] β-D-Galp 1.80 2JFE[X] 2.7 2ZOX[A] α-D-Glcp-(1-4)-GLU β-D-Glcp-(1-1)-<non_carb> 1.90 3VKK[A] α-D-Manp-(1-4)-GLU 2.00
β-Klotho (KLB;G6VTS68545; klotho-like)		Homo sapiens	BAC56857.1 AAH33021.1 AAI04872.1 AAI13654.1 AAI43576.1 AAQ41832.1 BAG63737.1 NP_783864.1		5VAN[A] D-1-deoxy-GlcpN D-1-deoxy-Manp α-D-Manp β-D-Glcp 2.20 5VAQ[A] 2.61 6NFJ[A,D] 3.19
β-Klotho (KLB;G6VTS68545; klotho-like)		Homo sapiens	BAC56857.1 AAI04872.1 AAI13654.1 AAI43576.1 AAQ41832.1 NP_783864.1		5VAK[A] 1.70 5VAN[A] D-1-deoxy-GlcpN D-1-deoxy-Manp α-D-Manp β-D-Glcp 2.20 5VAQ[A] 2.61 5WI9[A,B] 2.70 6NFJ[A,D] 3.19
β-glucosidase (Bgl4;Bglhi)	3.2.1.21	Humicola grisea var. thermoidea IFO9854 / Humicola insolens RP86	BAA74958.1	O93784	4MDO[A] 2.60 4MDP[A] β-D-Glcp 2.05
β-glucosidase (BGLN1)	3.2.1.21	Nannochloropsis oceanica	5YJ7_A		5YJ7[A,B,C,D] 1.61
β-glucosidase (Bgl;NkBgl;NkBG;G1NkBG)	3.2.1.21	Neotermes koshunensis	BAB91145.1	Q8T0W7	3AHZ[A] 1.34 3AI0[A] β-D-Glcp-(1-1)-<non_carb> 1.40 3VIF[A] β-D-Glcp 1.00 3VIG[A] <carb_like_NOJ> 0.99 3VIH[A] 1.03 3VII[A] 0.97 3VIJ[A] β-D-Glcp 1.03 3VIK[A] β-D-Glcp-(1-4)-β-D-Glcp 1.10 3VIL[A] β-D-Glcp-(1-1)-<non_carb> 1.15 3VIM[A] β-D-Glcp-(1-1)-<non_carb> 1.03 3VIN[A] β-D-Glcp-(1-1)-<non_carb> 1.13 3VIO[A] β-D-Glcp-(1-1)-<non_carb> 1.12 3VIP[A] β-D-Glcp-(1-1)-<non_carb> 1.28
β-mannosidase (Os7BGlu26)	3.2.1.25	Oryza sativa Indica Group	ACF35791.1	B5ABY0	4JHO[A] 2.21 4JIE[A] β-D-Manp 2.45 4RE2[A] <carb_like_MVL> 2.00 4RE3[A] <carb_like_GIM> 2.55 4RE4[A] <carb_like_IFM> 2.29
β-glucosidase (Os03g0212800;Os3bglu6)	3.2.1.21	Oryza sativa Japonica Group	AAN01354.1 BAF11272.1 BAG99682.1 BAS82933.1 BBQ09600.1	Q8L7J2	3GNO[A] 1.83 3GNP[A] β-D-Glcp-(1-1)-octyl 1.80 3GNR[A] α-D-Glcp2F-(1-4)-GLU 1.81 3WBA[A] α-D-Glcp-(1-4)-GLU 1.90 3WBE[A] α-D-Glcp-(1-4)-GLU 1.97
β-glucosidase (OsTAGG2;Os04g0474800;Os4bglu12) (salicylic acid glucoside-hydrolyzing)	3.2.1.21 3.2.1.58 3.2.1.74	Oryza sativa Japonica Group	CAE05483.2 BAF14984.1 BAG94781.1 BAS89674.1 BAS89675.1 BAS89676.1 CAE05489.1	A0A0P0WB96 A0A0P0WBI4 A0A0P0WBJ3 Q0JCF0 Q7XKV4	3PTK[A,B] 2.49 3PTM[A,B] α-D-Glcp2F-(1-4)-GLU 2.40 3PTQ[A,B] β-D-Glcp2F-(1-1)-<non_carb> 2.45
monolignol β-glucosidase (Os4bglu18;Os04g0513900)	3.2.1.21	Oryza sativa Japonica Group	CAE01911.1 BAF15219.1 BAS90064.1 CAE01910.2 CAE54546.1	Q0JBR7 Q7XSJ9 Q7XSK0	7D6A[A,B] 1.70 7D6B[A,B] α-D-Glcp β-D-Glcp 2.10
β-glucosidase (Bglu1;Os03g0703000;Os3bglu7)	3.2.1.21	Oryza sativa Japonica Group	CBD28469.1 AAA84906.2 AAA84906.3 AAS07254.1 AAS07255.1 AAT85322.1 AAX95519.1 AKS44051.1 BAF12927.1 BAG94472.1 BAS85953.1 BAS85954.1 CBD28462.1 CBD28475.1 CBG21207.1	A0A067Y2M1 A0A0P0W1U0 Q42975 Q75I93	2RGL[A,B] 2.20 2RGM[A,B] α-D-Glcp2F-(1-4)-GLU 1.55 3AHT[A,B] β-D-Glcp-(1-3)-β-D-Glcp 2.80 3AHV[A,B] α-D-Glcp2F-(1-4)-GLU 1.89 3F4V[A,B] 1.65 3F5I[A,B] α-D-Glcp2F-(1-4)-GLU 1.75 3F5J[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 1.95 3F5K[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 1.80 3F5L[A,B] β-D-Glcp-(1-3)-β-D-Glcp 1.37 3SCN[A,B] 2.20 3SCO[A,B] α-D-Glcp1F 1.95 3SCP[A,B] 2.10 3SCQ[A,B] α-D-Glcp1F 2.10 3SCR[A,B] 1.80 3SCS[A,B] α-D-Glcp1F 1.85 3SCT[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 1.60 3SCU[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 1.58 3SCV[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 2.11 3SCW[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 1.90 4QLJ[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 1.95 4QLK[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 1.83 4QLL[A,B] β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp-(1-4)-β-D-Glcp 1.85 7BZM[A,B] <carb_like_GIM> 2.30
β-glucosidase (Bgl1A)	3.2.1.21	Phanerochaete chrysosporium K-3	BAE87008.1	Q25BW5	2E3Z[A,B] 1.50 2E40[A,B] β-D-Glcp 1.90
strictosidine β-glucosidase (Sgr1;SG)	3.2.1.105	Rauvolfia serpentina	CAC83098.1 CBG21226.1	Q8GU20	2JF6[A,B] 2.82 2JF7[A,B] 2.48 3ZJ7[A,B] 2.50 3ZJ8[A,B] 3.01
raucaffricine O-β-glucosidase (RG)	3.2.1.125	Rauvolfia serpentina	AAF03675.1	Q9SPP9	3U57[A,B] β-D-Glcp-(1-1)-<non_carb> 2.43 3U5U[A,B] 2.20 3U5Y[A,B] β-D-Glcp-(1-1)-<non_carb> 2.30 3ZJ6[A,B] 2.40 4A3Y[A,B] 2.15 4ATD[A,B] 2.10 4ATL[A,B] β-D-Glcp 2.52 4EK7[A,B] β-D-Glcp 2.30
β-glucosidase (ScGlu) (plastidic)	3.2.1.21 3.2.1.182	Secale cereale	AAG00614.1 CAW42885.1 CAW48588.1 CBC08413.1 CBD21381.1 CBG21237.1	Q9FYS3	3AIU[A] 2.20 3AIV[A] 2.50 3AIW[A] α-D-Glcp2F-(1-4)-GLU 2.40
myrosinase	3.2.1.147	Sinapis alba	CAA42533.1	P29736 Q7SIB0	1DWA[M] 2.0 1DWF[M] 2.00 1DWG[M] L-1-deoxy-Gulp 2.0 1DWH[M] L-1-deoxy-Gulp 2.0 1DWI[M] L-1-deoxy-Gulp 2.0 1DWJ[M] 2.40 1E4M[M] 1.2 1E6Q[M] L-1-deoxy-Gulp <carb_like_NTZ> 1.35 1E6S[M] <carb_like_GOX> 1.35 1E6X[M] α-D-Glcp 1.6 1E70[M] D-1-deoxy-Glcp2F 1.65 1E71[M] 1.50 1E72[M] <carb_like_GOX> 1.6 1E73[M] α-D-Glcp2F-(1-4)-GLU 1.50 1MYR[A] 1.64 1W9B[M] β-D-GlcpNAc <carb_like_CGT> 1.7 1W9D[M] β-D-GlcpNAc 1.6 2MYR[A] α-D-Glcp2F-(1-4)-GLU 1.60 2WXD[M] 1.60
cyanogenic β-glucosidase / dhurrinase 1 (Dhr1)	3.2.1.21	Sorghum bicolor P721N	AAC49177.1 CAW42873.1 CAW48576.1 CBC08401.1 CBD21369.1 CBG21205.1	Q41290	1V02[A,B,C,D,F] 1.9 1V03[A] β-D-Glcp-(1-1)-<non_carb> 2.0
β-glycosidase (β-gly;Sfbgly)	3.2.1.21 3.2.1.23 3.2.1.38	Spodoptera frugiperda	AAC06038.1	O61594	5CG0[A,B,C,D,E,F] 2.09
β-glucosidase (ThBgl2; THAR02_02251)	3.2.1.21	Trichoderma harzianum T6776 / IOC-3844	KKP05610.1		5JBO[A] 1.97
β-glucosidase (ThBgl1; THAR02_05432)	3.2.1.21	Trichoderma harzianum T6776 / IOC-3844	KKP02477.1		5JBK[A,B] 2.59
β-1,4-glucosidase (Bgl2;ThBgl)	3.2.1.21	Trichoderma harzianum T7	ABN50090.1 ABN50091.1	A0A0F9ZQA8 A0A2T4AR08 A3FPG4	5BWF[A,B] 2.60 6EFU[A,B] 2.20
β-glucosidase (Cel1b) (Cel1B)	3.2.1.21	Trichoderma reesei QM6A / QM9414	AAP57758.1 ABE22339.1 CBG21221.1	G0RIF5 Q7Z9M2	6KHT[A,B,C] 3.31
β-glucosidase 2 (CEL1a;Bgl2;BGLII;BGL) (Cel1A)	3.2.1.21	Trichoderma reesei QM9414	BAA74959.1	O93785	3AHY[A,B,C,D] 1.63
β-glucosidase 2 (cyanogenic)	3.2.1.21	Trifolium repens	ABV54716.1 ABV54717.1 ABV54718.1 ABV54719.1 ABV54720.1 ABV54721.1 ABV54722.1 ABV54723.1 ABV54724.1 ABV54725.1 ABV54726.1 ABV54727.1 ABV54728.1 ABV54729.1 ABV54730.1 ABV54731.1 ABV54732.1 ABV54733.1 ABV54734.1 ABV54735.1 ABV54736.1 ABV54737.1 ABV54738.1 ABV54739.1 ABV54740.1 ABV54741.1 ABV54742.1 ABV54743.1 ABV54744.1 ABV54745.1 ABV54746.1 ABV54747.1 ABV54748.1 ABV54749.1 CAA40057.1	A8C6G0 A8C6H2 A8C6J3 A8C6K7 A8C6L1 A8C6M3 P26205	1CBG[A] 2.15
β-glucosidase (Glu1B;TaGlu1b)	3.2.1.21 3.2.1.182	Triticum aestivum	BAE92259.1 AEV76983.1	J7F1W6 Q1XH05	2DGA[A] 1.80 3AIQ[A] 1.90 3AIR[A] α-D-Glcp2F-(1-4)-GLU 2.00 3AIS[A] β-D-Glcp-(1-1)-<non_carb> 2.20
zeatin β-glucosidase (Glu1;p60.1;Zm-p60.1)	3.2.1.21	Zea mays	CAA52293.1 CAM55487.1 CAM55549.1 CAW42858.1 CAW42907.1 CAW48561.1 CAW48562.1 CAW48610.1 CBC08386.1 CBC08387.1 CBD21354.1 CBD21355.1	P49235 Q53WW9	1HXJ[A,B] 2.05
β-glucosidase 1 (Glu1)	3.2.1.21 3.2.1.182	Zea mays	AAA65946.1 AAB03266.1 AAD10503.1 ACF82863.1 ACG32200.1 CAM55450.1 CAM55655.1 CAW42888.1 CAW48591.1 CBC08416.1 CBD21384.1	P49235 Q53WW9	1E1E[A,B] 2.5 1E1F[A,B] β-D-Glcp-(1-1)-<non_carb> 2.6 1E4L[A,B] 2.2 1E4N[A,B] 2.1 1E55[A,B] β-D-Glcp-(1-1)-<non_carb> 2.0 1E56[A,B] β-D-Glcp-(1-1)-<non_carb> 2.1 1H49[A,B] β-D-Glcp-(1-1)-<non_carb> 1.9 1V08[A,B] <carb_like_NTZ> 1.9
unclassified
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D Carbohydrate Ligands Resolution (Å)
ancestral heme-binding TIM-barrel glycosidase		synthetic construct			6Z1H[A,B] 2.50 6Z1M[A,B,C] 2.45
β-glucosidase	3.2.1.21	uncultured microorganism	BAV69317.1	A0A1E1FFN6	5XGZ[A,B] 1.75 7WDN[A] α-D-Glcp 1.80 7WDO[A] β-D-Glcp β-D-Glcp-(1-4)-β-D-Glcp 2.21 7WDP[A] α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp 2.39 7WDR[A] β-D-Glcp-(1-1)-<non_carb> 2.00 7WDS[A] β-D-Xylp 1.68 7WDV[A] β-D-Glcp β-D-Glcp-(1-4)-β-D-Glcp 1.81
β-glucosidase (Ambgl-lP)	3.2.1.21	unidentified	SOU12973.1	A0A2I2LGB3	5WKA[A,B,C,D] 2.75
β-glucosidase BglM-G1 from marine metagenome		unidentified			5NS6[B,C] 1.50 5NS7[A,B,C] 1.54 5NS8[A,B,C] <carb_like_NOJ> 1.55 5NS6[A] 1.50
β-glucosidase (Td2F2)	3.2.1.21	unidentified			3WH5[A] 1.60 3WH7[A] β-D-Fucp 1.10 3WH8[A] <carb_like_IFM> 1.90 5AYB[A] 1.80 5AYI[A] β-D-Glcp 1.85 3WH6[A] α-D-Glcp β-D-Glcp 1.60
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