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Glycoside Hydrolase Family 32

Activities in FamilyLevan fructotransferase (DFA-IV-forming) (EC 4.2.2.16);
Mechanism Retaining a
ClanGH-J
3D Structure Status5-fold β-propeller
Catalytic Nucleophile/BaseAsp
Catalytic Proton DonorGlu
External resourcesCAZypedia; PROSITE;
Commercial Enzyme Provider(s)MEGAZYME;
Statistics GenBank accession (28143); Uniprot accession (299); PDB accession (85); 3D entries (25); cryst (2)
Summary
Download GH32 (29861) Taxonomic display Structure (25 - 2 cryst) Characterized (227) Activities in Family (14)
Bacteria
Protein Name EC#ReferenceOrganismGenBank UniprotPDB/3D
 invertase (LevJ) 3.2.1.26 pubmed
pubmed
 Actinomyces naeslundii T14V AAA67876.1 Q44109  
 invertase (LevJ) 3.2.1.26 pubmed
pubmed
 Actinomyces naeslundii T14V AAA67876.1 Q44109  
 β-fructosidase (GH32-1; EUBREC_0257) 3.2.1.26 pubmed
 Agathobacter rectalis ATCC 33656 ACR74061.1 C4ZAK0  
 β-fructosidase (GH32-3; ANBU17_15490) 3.2.1.26 pubmed
 Anaerostipes butyraticus JCM 17466 GFO85652.1 A0A916Q7K1  
 β-fructosidase (Scrb_2; GH32-1; ANBU17_09520) 3.2.1.26 pubmed
 Anaerostipes butyraticus JCM 17466 GFO84605.1 A0A916Q8H4  
 β-fructosidase (ScrB; ANCC_10700) 3.2.1.26 pubmed
 Anaerostipes caccae JCM 13470 BCD35034.1    
 β-fructosidase (GH32-1; DO83_03400) 3.2.1.26 pubmed
 Anaerostipes hadrus BPB5 AQP38736.1 A0A173T8W3  
 β-fructosidase (GH32-4; DO83_10515) 3.2.1.26 pubmed
 Anaerostipes hadrus BPB5 AQP39972.1 A0A1Q2C894  
 β-fructosidase (GH32-2; DO83_03965) 3.2.1.26 pubmed
 Anaerostipes hadrus BPB5 AQP38830.1 A0A1Q2C595  
 inulin exo-β-2,1-fructosidase (InuA) 3.2.1.153 pubmed
 Arthrobacter sp. MN8 AGC01505.1 A0A0F6MV26 8I12[A]
 endo-inulinase (Inu1) 3.2.1.7 pubmed
 Arthrobacter sp. S37 CAB63119.1 Q9RBA5  
 endolevanase (LevB1) (partial) 3.2.1.65 doi
 Bacillus licheniformis IBT-ALM / IBt1 AHK25455.1 W8GV60 9H8I[A]
 endo-levanase 3.2.1.65 pubmed
 Bacillus sp. L7 CAA73180.1 O31411  
 invertase (SacA;BSU38040;IPA-50D) 3.2.1.26 pubmed
 Bacillus subtilis subsp. subtilis str. 168 AAA22723.1
ABV89908.1
AIY95118.1
AQR83843.1
AQR88058.1
CAA51606.1
CAB15830.1
CAB15830.2
NP_391683.1		 A8HU42
P07819		  
 levanase (SacC; BSU27030) 3.2.1.65 pubmed
 Bacillus subtilis subsp. subtilis str. 168 CAB14645.1
AIY94027.1
AQR82724.1
AQR86939.1
CAA29137.1
CAA63465.1
CAA68542.1
NP_390581.1		 A0A6M3ZED6
P05656		  
 endo-levanase (YveB;LevB;BSU34460) 3.2.1.65 pubmed
 Bacillus subtilis subsp. subtilis str. 168 CAB08014.1
AIY94762.1
AQR83480.1
AQR87695.1
CAB15451.1
NP_391326.1		 A0A6M4JPE5
O07003		  
 β-fructanase (FruA;ScrL) 3.2.1.26
3.2.1.64
3.2.1.65		 pubmed
pubmed
pubmed
 Bacteroides fragilis BF-1 AAA22924.1 Q45155  
 β-fructofuranosidase (BT1759; BT_1759) 3.2.1.26
3.2.1.64
3.2.1.153		 pubmed
pubmed
pubmed
 Bacteroides thetaiotaomicron VPI-5482 AAO76866.1
NP_810672.1		 Q8A6W7  
 exo-β-fructofuranosidase (BT3082; BT_3082) 3.2.1.26
3.2.1.64
3.2.1.153		 pubmed
pubmed
pubmed
 Bacteroides thetaiotaomicron VPI-5482 AAO78188.1
NP_811994.1		 Q8A373  
 endo-β-2,6-fructanase / endo-levanase (BT1760; BT_1760) 3.2.1.65 pubmed
 Bacteroides thetaiotaomicron VPI-5482 AAO76867.1
NP_810673.1		 Q8A6W6 6R3R[A]
6R3U[A]
7ZNR[A,B]
7ZNS[A,B]
8A9Y[A,C]
8AA0[A,C]
8AA2[A,D]
 exo-β-fructofuranosidase (BT1765; BT_1765) 3.2.1.26
3.2.1.64		 pubmed
pubmed
 Bacteroides thetaiotaomicron VPI-5482 AAO76872.1
NP_810678.1		 Q8A6W1  
 β-fructofuranosidase (CscA) 3.2.1.26 pubmed
 Bifidobacterium adolescentis G1 BAJ21238.1 E1CPU4  
 β-fructosidase (BfrA) 3.2.1.26 pubmed
 Bifidobacterium animalis subsp. lactis DSM 10140(T) CAD26970.1
AAS87041.1		 Q564C7
Q84F22		  
 β-fructofuranosidase (FosC; Bbr_1324) 3.2.1.26 pubmed
 Bifidobacterium breve UCC2003 AAT28190.1
ABE95999.1		 F9Y237
Q4JEX8		  
 inulin exo-β-2,1-fructosidase (BBDE_1523) 3.2.1.153 pubmed
 Bifidobacterium dentium JCM 1195 = DSM 20436 BAQ27517.1    
 β-fructofuranosidase 3.2.1.26 doi
 Bifidobacterium longum KN29.1 ABN04092.1 A2TLS9 3PIG[A,B]
3PIJ[A,B]
 sucrose-6-P hydrolase (ScrB;SacA;CAC0425) 3.2.1.26 pubmed
 Clostridium acetobutylicum ATCC 824 AAF35841.1
AAK78405.1
NP_347065.1		 Q7D471
Q9L8G4		  
 invertase 3.2.1.26 pubmed
 Clostridium beijerinckii AAC99322.1 Q9ZHJ7  
 β-fructosidase (GH32-1; COEU31_05240) 3.2.1.26 pubmed
 Coprococcus eutactus JCM 31265 GFO93478.1 A0AAI9K369  
 β-fructosidase (SacA; GH32-2; COEU31_16540 3.2.1.26 pubmed
 Coprococcus eutactus JCM 31265 GFO94608.1 A0AAI9NYB8  
 sucrose-6-P hydrolase (ScrB;Cgl2643;Cg2643) 3.2.1.26 pubmed
 Corynebacterium glutamicum ATCC 13032 BAC00037.1
CAF21305.1
CCH25774.1
NP_601843.1		 Q8NMD5  
 sucrose hydrolase (CscA) 3.2.1.26 pubmed
 Escherichia coli AAP79503.1 O86076  
 invertase (CscA) 3.2.1.26 pubmed
 Escherichia coli B-62 AAC33123.1 O86076  
 invertase (CscA) 3.2.1.26 pubmed
 Escherichia coli EC3132 CAA57219.1 P40714  
 β-fructosidase (CG447_09260) 3.2.1.26 pubmed
 Faecalibacterium prausnitzii A2165 ATP01188.1    
 β-fructofuranosidase (FperFFase;FPB0191_01269) 3.2.1.26 pubmed
 Frischella perrara PEB0191 AJA45090.1
BCZ08027.1		 A0A0A7S0J1 7VCO[A]
7VCP[A]
 levanase (SurC) 3.2.1.65 pubmed
 Geobacillus stearothermophilus ATCC 12980 AAB97112.1 P94469  
 exo-inulinase (InuA) 3.2.1.153 pubmed
 Geobacillus stearothermophilus KP1289 BAC45010.1 Q8GI55 cryst
 invertase (SurA) 3.2.1.26 pubmed
 Geobacillus stearothermophilus NUB36 AAB38976.1 P94466  
 levanase (LsdB) 3.2.1.65 doi
 Gluconacetobacter diazotrophicus SRT4 CBS 550.94 AAF16405.1
QEP49750.1		 Q9RBJ1  
 invertase (ScrB) 3.2.1.26 pubmed
 Klebsiella pneumoniae 1033-5P14 CAA40659.1
CAA47976.1		 P27217  
 inulinase / fructan hydrolase (LevH1) 3.2.1.153 pubmed
 Lacticaseibacillus casei IAM 1045 BAD88632.1 Q5H7T0  
 β-fructosidase (FosE) 3.2.1.65 pubmed
 Lacticaseibacillus paracasei 1195 ABD57319.1 Q27J21  
 fructan β-fructosidase / inulinase (Inu; InuB41) 3.2.1.7 pubmed
 Lacticaseibacillus paracasei subsp. paracasei DSM 23505 AKA87712.1 A0A0E3KC98  
 β-fructofuranosidase (SacA;LPST_C0153) 3.2.1.26 doi
 Lactiplantibacillus plantarum ST-III ADN97377.1 E1TRE0  
 sucrose-6-phosphate hydrolase (LgAs32; LGAS_1779) 3.2.1.26 pubmed
 Lactobacillus gasseri ATCC 33323 = JCM 1131 ABJ61066.1 A0A806A0W7
D1YK18		 6NU7[A]
6NU8[A]
 invertase (ScrB;SacA) 3.2.1.26 pubmed
 Lactococcus lactis NIZO R5 AAA25218.1
CAB09689.1		 Q04937
Q797W8		  
 inulin exo-β-2,1-fructosidase (MfFEH; J0P97_13015) 3.2.1.153 pubmed
 Microbacterium flavum DSM 18909 MBT8798983.1    
 sucrose/inulin exo-β-2,1-fructosidase (MfFFase1; J0P97_08245) 3.2.1.26
3.2.1.153		 pubmed
pubmed
 Microbacterium flavum DSM 18909 MBT8798059.1    
 sucrose β-2,1-fructosidase (MfFFase2; J0P97_08250) 3.2.1.26 pubmed
 Microbacterium flavum DSM 18909 MBT8798060.1    
 2,6-β-fructan 6-levanbiohydrolase (LevM) 3.2.1.64 pubmed
 Microbacterium laevaniformans ATCC 15953 BAB59060.1 Q93R69  
 levan fructotransferase (LftM) 4.2.2.16   Microbacterium sp. AL-210 AAG01554.1 Q9EVQ9  
 β-fructofuranosidase (SacA; FF33) 3.2.1.26 pubmed
 Microbulbifer sp. JAM-3301 BAL70276.1
BAL70274.1		 H7CE75  
 endo-inulinase (Inu; IN33) 3.2.1.7 pubmed
 Microbulbifer sp. JAM-3301 BAL70278.1
BAL70275.1		 H7CE76  
 cycloinulo-oligosaccharide fructanotransferase 2.4.1.- pubmed
 Niallia circulans MCI2554 BAA24360.1 O52973  
 levan fructosyltransferase (Lft) 4.2.2.16   Paenarthrobacter nicotinovorans GS-9 BAA24915.1 O50585  
 levan fructotransferase (DFA-IV-forming) (LftA) 4.2.2.16   Paenarthrobacter ureafaciens K-2032 AAF73829.1
AAX01526.1
AAX01527.1		 Q9KJD0 4FFF[A,B,C,D]
4FFG[A,B,C,D]
4FFH[A,B,C,D]
4FFI[A,B,C,D]
 cycloinulo-oligosaccharide fructanotransferase (Cft) 2.4.1.- pubmed
 Paenibacillus macerans CFC1 AAG47946.1 Q9F0I5  
 levanase (LelA) 3.2.1.65 pubmed
 Paenibacillus polymyxa CF43 CAA81392.1 Q45372  
 cycloinulo-oligosaccharide fructanotransferase (CFTase) 2.4.1.- doi
 Paenibacillus polymyxa MG-CF6 BAM74653.1 L8AYK6  
 exo-inulinase (Inu3) 3.2.1.153 pubmed
 Paenibacillus polymyxa MGL21 AAL82575.1 Q8RLU2  
 cycloinulo-oligosaccharide fructanotransferase (Cft;CFTase) 2.4.1.- doi
 Paenibacillus polymyxa MGL21 AAL82574.1 Q8RLU3  
 exo-β-1,2-fructosidase / fructanase (FruA;BWX39_02900) 3.2.1.153 pubmed
 Prevotella intermedia ATCC 25611 = DSM 20706 APW31681.1
BAI39500.1		 A0A1P8JD73
C8KIP0		  
 β-fructosidase A (FruA) 3.2.1.26 pubmed
 Priestia megaterium ATCC 14581 AAM19071.1 Q8GM36  
 inulin endo-β-2,1-fructosidase (SacAP2; FruA; EQG57_07990) 3.2.1.7 pubmed
 Priestia megaterium NBRC 15308 = ATCC 14581 QCY24540.1    
 sucrose β-2,1-fructosidase (SacAP3; EQG57_19615) 3.2.1.26
2.4.1.-		 pubmed
pubmed
 Priestia megaterium NBRC 15308 = ATCC 14581 QCY26665.1    
 sucrose β-2,1-fructosidase (SacAP1; EQG57_20070) 3.2.1.26 pubmed
 Priestia megaterium NBRC 15308 = ATCC 14581 QCY26750.1    
 endo-inulinase (Inu1) 3.2.1.7 URL
 Pseudomonas mucidolens AAF24999.1 Q9RGC0  
 exo-inulinase (Inu2) 3.2.1.153 URL
 Pseudomonas mucidolens AAF44125.1 Q9LAL1  
 β-fructosidase (M72_24121) 3.2.1.26 pubmed
 Roseburia faecis M72 CRL35568.1 A0A0M6WGE1  
 β-fructosidase (SacC; RIL182_04003) 3.2.1.26 pubmed
 Roseburia intestinalis L1-82 VCV24083.1 C7G9C0  
 β-fructosidase 3.2.1.26 pubmed
 Roseburia inulinivorans DSM 16841 A2-194 ADF57367.1 C0FYJ5  
 invertase (ScrB) 3.2.1.26 pubmed
 Salmonella typhimurium CAA47974.1 P37075  
 exo-inulinase (InuA;InuAGN25) 3.2.1.153 doi
 Sphingobacterium sp. GN25 AGC01503.1 A0A0E3D7Z1  
 inulinase (InuA;InuAJB13) 3.2.1.153 pubmed
 Sphingomonas sp. JB13 AFE82292.1 A0A0F6MTX2  
 invertase (ScrB) 3.2.1.26 pubmed
 Staphylococcus xylosus CAA47970.1 Q05936  
 fructan β-fructosidase (FruA) 3.2.1.7
3.2.1.65		 pubmed
pubmed
 Streptococcus mutans GS-5 AAA26889.1 Q03174  
 invertase (ScrB) 3.2.1.26 pubmed
 Streptococcus mutans GS-5 AAA26972.1
CAA35872.1		 P13522  
 β-fructosidase (SusH;SP1795;SP_1795) 3.2.1.26 pubmed
 Streptococcus pneumoniae TIGR4 AAK75868.1
NP_346228.1		 A0A0H2URL9
Q97P49		  
 sucrose-6-P hydrolase (ScrH;SP1724) 3.2.1.26 pubmed
 Streptococcus pneumoniae TIGR4 AAK75801.1
NP_346161.1		 A0A0H2URG2
Q97PB6		  
 sucrose-6-phosphate hydrolase (ScrB) 3.2.1.26 pubmed
 Streptococcus sobrinus S68598    
 β-2,6-fructan 6-levanbiohydrolase (Lf2) 3.2.1.64 pubmed
 Streptomyces exfoliatus F3-2 BAC54106.1 Q8GI79  
 invertase (BfrA;Tm1414) 3.2.1.26 pubmed
 Thermotoga maritima MSB8 AAD36485.1
AGL50344.1
AHD18692.1
AKE29174.1
CAA04518.1
NP_229215.1		 G4FFE9
O33833		 1UTW[A,B,C,D,E,F]
1UYP[A,B,C,D,E,F]
1W2T[A,B,C,D,E,F]
 invertase (Inv) 3.2.1.26 doi
 uncultured bacterium ACN59531.1 C0LIF8  
 invertase (Inv2;UnInv2) 3.2.1.26 pubmed
 uncultured bacterium ADZ31185.1 F1CGX0  
 β-fructofuranosidase (MetaPbio_Contig10a_40--FOS) 3.2.1.26 pubmed
 uncultured bacterium CCG34932.1 S6CPR8  
 β-fructofuranosidase (MetaPbio_contig9a_30c--FOS) 3.2.1.26 pubmed
 uncultured bacterium CCG34775.1 S6CPG6  
 invertase (ScrB) 3.2.1.26 pubmed
 Vibrio alginolyticus AAA27562.1 P13394  
 invertase A (SacA) 3.2.1.26 pubmed
 Zymomonas mobilis AAA27701.1 F8DVG5
P22632
P35636		  
 invertase (InvA;ZMO0942) 3.2.1.26 pubmed
 Zymomonas mobilis subsp. mobilis ZM4 = ATCC 31821 AAV89566.1
AVZ42197.1		 P0DJA7  
 invertase E1 (InvA) 3.2.1.26 pubmed
 Zymomonas mobilis Z6C BAA01258.1 P0DJA7
P35636
P35636		  
Eukaryota
Protein Name EC#ReferenceOrganismGenBank UniprotPDB/3D
 sucrose:sucrose 1-fructosyltransferase (Sst2; Atq1-SST2; Atq1-SST-2) 2.4.1.99 pubmed
 Agave tequilana AFJ21577.1
AFJ21571.1		 I1ZBQ5  
 fructan:fructan 6G-fructosyltransferase (Fft1) 2.4.1.243 pubmed
 Agave tequilana AFJ21572.1 I1ZBQ0  
 cell wall invertase (Cwinv1; AtqCwinv-1) 3.2.1.26 pubmed
 Agave tequilana AFJ21574.1 I1ZBQ2  
 fructan:fructan 6G-fructosyltransferase (Fft1; Atq6G-FFT1) 2.4.1.243 pubmed
 Agave tequilana AFJ21578.1 I1ZBQ6  
 sucrose:sucrose 1-fructosyltransferase (Sst1;1-SST) 2.4.1.99 doi
 Agave tequilana ABG00265.1 A7RDD3  
 sucrose:sucrose 1-fructosyltransferase 2.4.1.99 pubmed
 Allium cepa CAA03099.1    
 invertase (Inv) 3.2.1.26 pubmed
 Allium cepa CAA06839.1 O81083  
 sucrose:sucrose 1-fructosyltransferase (SST-1;1-SST) 2.4.1.99 pubmed
 Allium cepa CAA06838.1 O81082  
 fructan:fructan 6G-fructosyltransferase (6-G FFT;6G-FFT) 2.4.1.243 pubmed
 Allium cepa CAA69170.1 P92916  
 β-fructofuranosidase (SucA) 3.2.1.26 pubmed
 Amylomyces rouxii CBS 438.76 BAM95319.1 M1VKR3  
 invertase (AtBFruct4;Fruct4;AtbFruct4;At1g12240) 3.2.1.26 pubmed
 Arabidopsis thaliana AAG12569.1
AAK76683.1
AAL65639.1
AAL65640.1
AAL65641.1
AAL65642.1
AAL65643.1
AAL65644.1
AAL65645.1
AAL65646.1
AAL65647.1
AAL65648.1
AAL65649.1
AAL65650.1
AAL65651.1
AAL65652.1
AAL65653.1
AAL65654.1
AAL65655.1
AAL65656.1
AAL65657.1
AAL65658.1
AAL65659.1
AAN13204.1
AEE28855.1
CAA66330.1
CAA72321.1
NP_563901.1		 Q39041
Q7DLW9
Q8L3Z1
Q8L488
Q8L4G6
Q8L4T3
Q8L4U9
Q8L514
Q8LGP2		  
 At1g62660 (invertase;AtBFruct3;Fruct3;AtcwINV2) 3.2.1.26 pubmed
 Arabidopsis thaliana AAN18078.1
AAD43622.1
AAF19535.1
AAK62665.1
AAK82531.1
AAL32559.1
AAL36260.1
AAM45114.1
AEE33991.1
ANM61060.1
ANM61061.1
ANM61062.1
CAA61624.1
CAA64781.1
CAA67560.1
NP_176453.1
NP_564798.1		 A0A1P8AWL7
A0A1P8AWQ1
Q42567
Q43348
Q94BX0
Q9SI83
Q9SXD2		  
 cell-wall invertase 1 (AtBFruct1;Fruct1;AtcwINV1;At3g13790) 3.2.1.26 pubmed
 Arabidopsis thaliana AAK76450.1
AAL85153.1
AEE75414.1
AEE75415.1
ANM65335.1
ANM65336.1
ANM65337.1
BAB01930.1
BAH56786.1
CAA52619.1
CAA52620.1
NP_187995.1
NP_566464.1		 A0A1I9LRX7
A0A5S9XBW4
C0Z257
F4JEJ0
Q43866
Q9LIB8		 2AC1[A]
2OXB[A]
2QQU[A]
2QQV[A]
2QQW[A]
2XQR[A,C,E,G,I,K]
 invertase 4 (Atfruct6;AtcwINV4;At2g36190) 3.2.1.26 pubmed
 Arabidopsis thaliana AAD21446.2
AAM15406.1
AAM61359.1
AEC09214.1
BAB83031.1
NP_565837.1		 Q8L5V2
Q8W413		  
 fructan 1-exohydrolase (AlEH2) 3.2.1.153 doi
 Arctium lappa BAR88557.1 A0A0H5AH66  
 fructan:fructan 1-fructosyltransferase (AlFt1;1-FEH) 2.4.1.100 doi
 Arctium lappa BAJ24841.1 E2RV71  
 fructan 1-exohydrolase (AlEH1;1-FEH) 3.2.1.153 pubmed
 Arctium lappa BAL73222.1 I0FXI5  
 fructan:fructan 6G-fructosyltransferase (AoFT1;6-G FFT;6G-FFT) 2.4.1.243 pubmed
 Asparagus officinalis BAD89564.1 Q5FC15  
 exo-inulinase (Inu1) 3.2.1.153 pubmed
 Aspergillus awamori var. 2250 CAC44220.1 Q96TU3 1Y4W[A]
1Y9G[A]
1Y9M[A]
 endo-inulinase (Inu2) 3.2.1.7 pubmed
 Aspergillus ficuum ATCC 16882 CAA07345.1
ACR55693.1		 C5J3N4
O94220		 3RWK[X]
3SC7[X]
 sucrose:sucrose 1-fructosyltransferase (1-Sst) 2.4.1.99 pubmed
 Aspergillus foetidus CAA04131.1 O42801  
 β-fructofuranosidase (FopA) 3.2.1.26 pubmed
 Aspergillus japonicus ATCC 20611 BAB67771.1 Q96VC5  
 fructosyltransferase (FT) 2.4.1.100 pubmed
 Aspergillus japonicus CB05 ADK46938.1 F1ADK9
Q1W3Z7		 3LDK[A]
3LDR[A]
3LEM[A]
3LF7[A]
3LFI[A,B]
3LIG[A]
3LIH[A]
 fructosyltransferase (AKAW_06215) 2.4.1.100 pubmed
 Aspergillus luchuensis IFO 4308 GAA88101.1 G7XM46 5XH8[A]
5XH9[A]
5XHA[A]
 exo-inulinase (InuE) 3.2.1.153 pubmed
 Aspergillus niger 12 BAD01476.1 Q76HP6 cryst
 endo-inulinase A (InuA) 3.2.1.7 pubmed
 Aspergillus niger 12 BAA33797.1 O74641  
 endo-inulinase B (InuB) 3.2.1.7 pubmed
 Aspergillus niger 12 BAA33798.1 O74642  
 endoinulinase (EndoInu) 3.2.1.7 URL
 Aspergillus niger 9891 AAN64131.1
AAO18341.1		 Q874R2
Q8J229		  
 inulinase (InuA1) 3.2.1.7 doi
 Aspergillus niger AF10 AAK43726.1 O74641
Q96WZ8		  
 β-fructofuranosidase (Suc1) 3.2.1.26 pubmed
 Aspergillus niger B60 S33920    
 invertase (SucA;Suc1;An08g11070;ANI_1_2508074) 3.2.1.26 pubmed
 Aspergillus niger CBS 513.88 ABB59678.1
CAK45778.1		 A2QSK6
Q0ZR37		  
 invertase (SucB; An15g00320) (intracellular) 3.2.1.26 pubmed
 Aspergillus niger CBS 513.88 ABB59679.1
CAK42201.1		 A2R4G3
Q0ZR36		  
 exo-inulinase E (InuE;Inu1;An12g08280) 3.2.1.153
2.4.1.99		 doi
doi
 Aspergillus niger CBS 513.88 ABB59682.1
CAK41278.1		 A2R0E0  
 invertase (Ifv) 3.2.1.26 doi
 Aspergillus niger GH1 ADR80690.1
AFU93429.1		 E5LLD8
K4K2F9		  
 invertase (Sir-1) 3.2.1.26 doi
 Aspergillus niger IBT10sb AAC08047.1 O13388  
 fructosyltransferase (Fts1) 2.4.1.100 pubmed
 Aspergillus niger QU10 AHC54391.1 V9SEC7  
 β-fructosidase (BfrA) / levan and- neolevan- β-2,6-fructosyltransferas 3.2.1.26
2.4.1.-		 pubmed
pubmed
 Aspergillus oryzae FS4 AHY94898.1
AHY94899.1		 A0A024A3C6  
 fructosyltransferase (AoFT;AO090701000400) 2.4.1.- pubmed
 Aspergillus oryzae RIB40 BAE62112.1 Q2U8K3  
 2,1-fructan:2,1-fructan 1-fructosyltransferase (Sft;SftA) 2.4.1.100 pubmed
 Aspergillus sydowii IAM 2544 CAB89083.1
AAY07202.1
ACW62827.1		 Q9P853  
 invertase (cell-wall) (BObetaFRUCT1) 3.2.1.26 pubmed
 Bambusa oldhamii ABB77249.1 Q2XQ21  
 invertase (cell-wall) (BObetaFRUCT2) 3.2.1.26 pubmed
 Bambusa oldhamii ABB77250.1 Q2XQ20  
 invertase (vacuolar) (BObetaFRUCT3) 3.2.1.26 pubmed
 Bambusa oldhamii ABB77251.1 Q2XQ19  
 fructan b-(2,6)-fructosidase (Fru-1;6-FEH) 3.2.1.154 pubmed
 Beta vulgaris CAD48404.1 Q70XE6  
 β-fructofuranosidase (invertase; Suc1) 3.2.1.26 pubmed
 Bombyx mori BAG30882.1 B2DD57  
 β-fructofuranosidase (BmSUC1) 3.2.1.26 pubmed
 Bombyx mori Fuyo x Tsukubane BCD57653.1 A0A6F8Z6Y2 7BWB[A]
7BWC[A]
 acid invertase (AI7-3) 3.2.1.26 doi
 Brassica oleracea AAG36943.1
VDD42296.1		 A0A3P6EXQ9
Q9FQ61		  
 acid invertase (AI6-1) 3.2.1.26 doi
 Brassica oleracea AAG36942.1
VDD59064.1		 A0A3P6GKC0
Q9FQ62		  
 fructan 1-exohydrolase (1-FEH) 3.2.1.153 pubmed
 Bromus pictus ACZ65470.1 D2IGW7  
 β-fructofuranosidase (Inv;CmINV) 3.2.1.26 pubmed
 Ceratocystis moniliformis CMW 10134 AGV22100.1 T2C607  
 fructan fructan 1-fructosyltransferase (1-FFT) 2.4.1.100 pubmed
 Cichorium intybus AAD00558.1
AAS23663.1
CAB70428.1
CAC07564.1		 Q9ZR96  
 invertase 3.2.1.26 pubmed
 Cichorium intybus CAA72062.1
AFU54057.1		 Q9ZR54
V9LTM5		  
 sucrose:sucrose 1-fructosyl transferase (1-SST) 2.4.1.99 pubmed
 Cichorium intybus AAB58909.1
CAB70429.1		 O24459  
 fructan b-(2,1)-fructosidase / fructan 1-exohydrolase I (1-FEH I) 3.2.1.153 pubmed
 Cichorium intybus CAC19366.1
AFU54054.1
AFU54055.1
AIP90172.1		 A0A0K0KU50
Q9FNS9
V9LTH3
V9LTP0		  
 fructan 1-exohydrolase IIb (1-FEH IIb) 3.2.1.7 pubmed
 Cichorium intybus CAC37923.1
AFU54058.1
AFU54059.1
AIP90174.1		 A0A0K0KUJ3
Q93X59
V9LTI5
V9LTP1		  
 fructan b-(2,1)-fructosidase; fructan 1-exohydrolase IIa (1-FEH IIa) 3.2.1.153 pubmed
 Cichorium intybus CAC37922.1
AAP85536.1
AFU54056.1		 Q93X60
V9LTK7		 1ST8[A]
2ADD[A]
2ADE[A]
2AEY[A]
2AEZ[A]
 invertase (Inv1) 3.2.1.26 pubmed
 Cyberlindnera jadinii NRRL-Y1084 CAA73208.1 O94224  
 fructan fructan 1-fructosyltransferase (1-FFT) 2.4.1.100 pubmed
 Cynara cardunculus var. scolymus CAA04120.2
AAP68061.1
AAP68062.1
ABI11210.1
ABI11211.1
CAB70496.1
CAB70497.1		 O65778  
 sucrose:sucrose 1-fructosyltransferase (1-SST) 2.4.1.99 pubmed
 Cynara cardunculus var. scolymus CAA70855.1
CAB69364.1
CAB69365.1		 O23786  
 invertase 1 (Inv1;Inva1*DC1) 3.2.1.26 pubmed
 Daucus carota AAA03516.1
CAA49162.1		 P26792  
 invertase (isoform I;Inv*DC4 ) 3.2.1.26 pubmed
 Daucus carota CAA53098.1
CAA53097.1
CAA77267.1		 P80065  
 invertase 3.2.1.26 pubmed
 Daucus carota CAA53099.1 P80065  
 invertase (Inv3;Inv*Dc3) (insoluble) 3.2.1.26 pubmed
 Daucus carota CAA55188.1 Q39693  
 invertase (isoform II;Inv*Dc5) 3.2.1.26 pubmed
 Daucus carota CAA47636.1
CAA77266.1		 Q42722  
 invertase 4 (Inv*Dc4') 3.2.1.26 pubmed
 Daucus carota CAA77268.1 Q9ZR32  
 2,1-fructan:2,1-fructan 1-fructosyltransferase (FFT) 2.4.1.100 pubmed
 Echinops ritro CAH18891.1 Q0PCC9  
 invertase (Suc2;AgSUC2;AFR529W) 3.2.1.26 pubmed
 Eremothecium gossypii ATCC 10895 AAS53900.1 Q752P4  
 1,2-β-fructan 1F-fructosyltransferase (FFT-1;1-FFT) 2.4.1.100 pubmed
 Helianthus tuberosus AAW04794.1
CAA03405.1
CAA08811.1		 O81985  
 sucrose:sucrose 1-fructosyltransferase (1-SST) 2.4.1.99 pubmed
 Helianthus tuberosus AAW04795.1
CAA03404.1
CAA08812.1		 O81986  
 sucrose:fructan 6-fructosyltransferase (6-SFT) 2.4.1.10 pubmed
 Hordeum vulgare AAW04797.1
CAA58235.1		 Q96466  
 soluble acid invertase (Inv1;HvInv1;Sai) 3.2.1.26 doi
 Hordeum vulgare CAF22241.1 Q6KCH6  
 apoplastic invertase 1 (CwInw1) 3.2.1.26 pubmed
 Hordeum vulgare subsp. vulgare CAD58960.1 Q8GT50  
 sucrose:sucrose 1-fructosyltransferase (1-SST) 2.4.1.99 doi
 Hordeum vulgare subsp. vulgare CAD98793.2 Q70LF5  
 invertase 1 (IbbFRUCT1;Ib2FRUCT1) 3.2.1.26 pubmed
 Ipomoea batatas AAD01606.1 Q9ZTX2  
 soluble acid invertase 3 (Ibβ-fruct3;FRUCT3) 3.2.1.26 pubmed
 Ipomoea batatas AAK71505.2 Q94C05  
 soluble acid invertase 2 (Ibainv1;Ibβ-Fruct2;Ibbetafruct2;FRUCT2) 3.2.1.26 pubmed
 Ipomoea batatas AAK71504.1
BAM74659.1		 L8AYN8
Q94C06		  
 inulinase (Inu1) 3.2.1.7 pubmed
 Kluyveromyces marxianus CBS 6556 CAA48500.1
CAA02437.1		 B3GQU5
P28999		  
 inulinase (Inu1;KcINU1;KcInu;KmInu1) 3.2.1.7 pubmed
 Kluyveromyces marxianus CBS4857 / DMKU3-1042 AAN32611.1
BAK08619.1		 Q8J2P7  
 exoinulinase 3.2.1.153 pubmed
 Kluyveromyces marxianus IW9801 CGMCC0360 AAT70412.1 Q6DQX2  
 inulinase (Inu1) 3.2.1.7 pubmed
 Kluyveromyces marxianus var. marxianus ATCC 12424 CAA40488.1
ACD93461.1		 B3GQU5
P28999
P28999		  
 invertase (CCW-inv) 3.2.1.26 pubmed
 Kummerowia stipulacea AFD62258.1 H9BP12  
 invertase (NCV-inv) 3.2.1.26 pubmed
 Kummerowia stipulacea AFD62255.1 H9BP09  
 invertase (CV-inv) 3.2.1.26 pubmed
 Kummerowia stipulacea AFD62256.1 H9BP10  
 invertase (NCW-inv) 3.2.1.26 pubmed
 Kummerowia stipulacea AFD62257.1 H9BP11  
 sucrose:sucrose 1-fructosyltransferase (1-SST) 2.4.1.99 pubmed
 Lolium arundinaceum CAC05261.1 Q9FSV7  
 fructan: fructan 6G-fructosyltransferase / sucrose:fructan 6-fructosyltransferase (6G-FFT;6-FT;1-FFT;6-SFT;Lp6-SFT) 2.4.1.10
2.4.1.243		 pubmed
pubmed
 Lolium perenne AAM14603.1
ABD72593.1
ABN25735.1
BAD35132.1
CAL51274.1		 Q05G11
Q6AW26
Q8S311		  
 fructan:fructan 6G-fructosyltransferase (6G-FFT;SSTLp6G-FFT_1;Prf3) / fructan:fructan 1F-fructosyltransferase (1-FFT) 2.4.1.100
2.4.1.243		 pubmed
pubmed
 Lolium perenne AAM13671.1
BAD26613.1
CAL51272.1		 Q05G13
Q6F4N3
Q8S337		  
 sucrose:sucrose 1-fructosyltransferase (1-SST;FT3;Lp1-SST) 2.4.1.99 pubmed
 Lolium perenne AAO86693.1
ABD72592.1
BAF99807.1
CAL51273.1		 A3QRG0
B0I1Q6
Q05G12
Q84RM0		  
 fructan exohydrolase 2 (1-FEHa; Lp1-FEHa) 3.2.1.153 pubmed
 Lolium perenne AAY81958.1
AAY81958.2		 Q108T5  
 exo-inulinase (Inu1) 3.2.1.153 pubmed
 Meyerozyma guilliermondii OUC1 ABW70125.2 A5DHM6
A8W7I5		  
 invertase (NCU04265) 3.2.1.26 pubmed
 Neurospora crassa OR74A CAC28747.2
XP_323605.1		 Q1K7I2
Q9C214		  
 Os04g0535600 (Vin1;OsVin1) 3.2.1.26 pubmed
 Oryza sativa Japonica Group CAD41518.1
BAF15336.1
BAG89376.1
BAS90254.1
BAS90255.1
CAD41525.3
CAE04784.2
CAE04785.1		 A0A0P0WCW5
A0A0P0WD12
Q0JBF1
Q7XM59
Q7XU69		  
 Os02g0106100 (Vin2;OsVin2) 3.2.1.26 pubmed
 Oryza sativa Japonica Group BAD28087.1
BAF07540.1
BAS76566.1		 Q0E4P8
Q6ETD3		  
 invertase (Cin1) 3.2.1.26 pubmed
 Oxybasis rubra CAA57389.1 Q42691  
 sucrose:{sucrose/fructan} 6-fructosyltransferase (6-Sst/6-Sft) 2.4.1.10
2.4.1.-		 pubmed
pubmed
 Pachysandra terminalis CBM41476.2
CBM41476.1		 E3PQS3 3UGF[A,B]
3UGG[A,B]
3UGH[A,B]
 exo-inulinase 3.2.1.153 pubmed
 Papiliotrema aurea G7a ACC61059.1 B2LYF0  
 exo-inulinase (Inu1) 3.2.1.153 pubmed
 Papiliotrema aurea HYA AFO84001.1 I7EDT5  
 exoinulinase (InuA1) 3.2.1.153 pubmed
 Penicillium janthinellum B01 AEO12134.1 G3GLJ6  
 β-fructofuranosidase (PoFF32A) 3.2.1.26 doi
 Penicillium oxalicum GXU20 AJY58099.1 A0A0D5MCQ0  
 exo-inulinase D (InuD; P-I) 3.2.1.153 pubmed
 Penicillium sp. TN-88 BAC16218.1 Q8J0G1  
 exo-inulinase D (InuD; P-I) 3.2.1.153 pubmed
 Penicillium sp. TN-88 BAC16218.1 Q8J0G1  
 endo-inulinase C (InuC) 3.2.1.7 pubmed
 Penicillium sp. TN-88 BAB19132.1 Q9HFA5  
 sucrose:{fructan/sucrose} 6-fructosyltransferase (PpFT1;6-Sft;6-Sst) 2.4.1.10
2.4.1.-		 pubmed
pubmed
 Phleum pratense BAH30252.1 C0SSP1  
 fructan 6-exohydrolase (6-FEH;Pp6-FEH1) 3.2.1.154 pubmed
 Phleum pratense BAJ76715.1 E9RGV6  
 invertase (PsB) 3.2.1.26 pubmed
 Pisum sativum CAA52143.1 Q43818  
 sucrose:{sucrose/fructan} 6-fructosyltransferase (6-SST;6-SFT) 2.4.1.10
2.4.1.-		 doi
doi
 Poa secunda AAG36767.1 Q9FR47  
 β-fructofuranosidase (SucA) 3.2.1.26 pubmed
 Rhizopus delemar NBRC 4754 BAM95316.1 M1V4E1  
 β-fructofuranosidase (Inv; RdINV) 3.2.1.26
2.4.1.99		 pubmed
doi
 Rhodotorula dairenensis CECT 1416 QFG75712.1 A0A856TAI5 8BEQ[A,B]
8BES[A,B,C,D]
8BET[A,B,C,D]
8BEU[A,B,C,D]
 invertase 2 (Suc2;YIL162w) 3.2.1.26 pubmed
 Saccharomyces cerevisiae S288c CAA24618.1
BAJ07819.1
CAA87030.1
NP_012104.1		 D5MS22
P00724		 4EQV[A,B,C,D,E,F,G,H]
 invertase (Inv1;SPCC191.11) 3.2.1.26 pubmed
 Schizosaccharomyces pombe 972h- CAD44470.1
AAE51667.1
BAA25684.1
CAB41057.1
NP_588300.1		 O59852  
 invertase / β-fructofuranosidase (Inv;SoINV;Ffase) 3.2.1.26 pubmed
 Schwanniomyces occidentalis ATCC 26077 ADN34605.1
AAE51668.1
CAA35606.1		 E5D0X5
P24133		 3KF3[A,B]
3KF5[A,B]
3U14[A,B]
3U75[A,B,C,D]
6S1T[A,B]
6S2B[A,B]
 β-fructofuranosidase 3.2.1.26 doi
 Scopulariopsis brevicaulis IFO4843 AAQ31829.1    
 invertase (Tiv1;TIV-1) 3.2.1.26 pubmed
 Solanum lycopersicum AAA34132.1
AAB30874.1
AAL75449.1
AAL75450.1
AAL99550.1
ASK06213.1
ASK06214.1
BAA01954.1
CAA78060.1
CAA78061.1
CAA78062.1
CAA78063.1		 A0A220QMI6
P29000
Q41215
Q547Q0
Q8RVW4
Q8S399		  
 invertase (Lin6;LIN6;Wiv-1) 3.2.1.26 pubmed
 Solanum lycopersicum AAM22409.1
AAM28823.1
AAV35206.1
BAA33150.1
CAA62735.1		 O82119
P93200
Q5QEW9
Q8LRN7
Q8LRN8		  
 acylsucrose fructofuranosidase 1 (Asff1; SnASFF1) 3.2.1.26 pubmed
 Solanum nigrum WDV74297.1    
 acylsucrose fructofuranosidase 1 (SpASFF1; Sopen03g040490) 3.2.1.26 pubmed
 Solanum pennellii QCH04025.1 A0A4D6U9U5  
 invertase (PAIN-1) 3.2.1.26 pubmed
 Solanum tuberosum CAA49831.1 Q43186  
 invertase (cwINV1;StcwINV1) 3.2.1.26 pubmed
 Solanum tuberosum CAA79676.1
AEV46321.1
AEV46322.1
AEV46323.1
AEV46324.1
AEV46325.1
CAA65086.1		 G9IHH8
G9IHH9
G9IHI0
G9IHI1
G9IHI2
Q43172		  
 β-fructofuranosidase (Sl-β-fruct) 3.2.1.26 pubmed
 Sphenophorus levis AIL92341.1 A0A088DJ59  
 endo-inulinase 3.2.1.7 pubmed
 Talaromyces purpureogenus HOK-01 BAA12320.1
BAA12321.1		 O00056  
 inulinase 3.2.1.153 pubmed
 Tanozyma kutaoensis CCF77887.1 H6SHQ3  
 sucrose:sucrose 1-fructosytransferase (Wft2;1-SST) 2.4.1.99 pubmed
 Triticum aestivum BAB82470.1
AAV26678.1
AMZ79593.1
AMZ79597.1
BAD72792.1		 A0A161H5V1
Q8W430		  
 sucrose:fructan 6-fructosyltransferase (Wft1;6-SFT) 2.4.1.10 pubmed
 Triticum aestivum BAB82469.1
AAV26677.1		 A0A9R1KAC3
Q8W431		  
 acid β-fructofuranosidase (Inv1;Vin2;TaVin2) 3.2.1.26 pubmed
 Triticum aestivum CAG25609.1 Q575T1  
 6-kesose β-2,6-fructosidase (A211;6-KEH w2) 3.2.1.- pubmed
 Triticum aestivum BAD99104.1 Q4W8R1  
 6-kesose β-2,6-fructosidase (C311;6-KEH w1) 3.2.1.- pubmed
 Triticum aestivum BAD99105.1 Q4W8R0  
 fructan:fructan 1-fructosyltransferase (1-FFT-A;Wft3) 2.4.1.100 pubmed
 Triticum aestivum BAE19751.1 Q4AEI9  
 fructan 6&1-exohydrolase (D31;6&1-FEH) 3.2.1.153
3.2.1.154		 pubmed
pubmed
 Triticum aestivum BAE44509.1 Q3MV21  
 fructan 6-exohydrolase (6-FEH) 3.2.1.154 pubmed
 Triticum aestivum CAJ28591.1 Q2UXF7  
 fructan 1-exohydrolase w3 (1-FEH w3) 3.2.1.153 pubmed
 Triticum aestivum PAJERO CAD92365.1 B6DZC8
Q70LR2		  
 invertase (Exin1) 3.2.1.26 pubmed
 Triticum monococcum AAN80141.1 Q8GT63  
 invertase 1 (Uf-Inv1) 3.2.1.26 pubmed
 Uromyces viciae-fabae CAG26671.1 Q2PCS4  
 fructan 1-exohydrolase (1-FEH) 3.2.1.153 pubmed
 Vernonia herbacea CAJ77148.1 A9JIF3  
 2,1-fructan:2,1-fructan 1-fructosyltransferase (Fft) 2.4.1.100 doi
 Viguiera discolor CAH18892.1 Q2WEC6  
 invertase (Inv1) 3.2.1.26 pubmed
 Wickerhamomyces anomalus CBS5759, ATCC8168 CAA56684.1 P40912  
 β-fructofuranosidase (Inv; Xd-INV; XdINV) 3.2.1.26 pubmed
 Xanthophyllomyces dendrorhous ATCC MYA-131 ACL79833.1 B8YJM2
J7HDY4		 5ANN[A,B]
5FIX[A,B]
5FK7[A,B]
5FK8[A,B]
5FKB[A,B]
5FKC[A,B]
5FMB[A,B]
5FMC[A,B]
5FMD[A,B]
5NSL[A,B]
5O47[A,B]
6FJE[A,B]
6FJG[A,B]
6S2G[A,B]
6S2H[A,B]
6S3Z[A,B]
6S82[A,B]
 invertase (Incw2;Mn1;Miniature1) 3.2.1.26 pubmed
 Zea mays AAF06991.1
AAC28320.1
AAD02510.1
ACG47298.1		 B6UD65
O81189
Q9SBI2
Q9SPK0		  
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