Auxiliary Activity Family 9

Mechanism monooxygenase
NoteAA9 (formerly GH61). The enzymes in this family were originally classified as a glycoside hydrolases (GH61) based on very weak endo-1,4-b-D-glucanase activity in one family member. They are now reclassified in the AA category of CAZy. Because a significant literature is associated with the old name GH61, we recommend to describe these enzymes as "AA9 (formerly GH61)".
External resourcesCAZypedia;
Statistics GenBank accession (1418); Uniprot accession (49); PDB accession (108); 3D entries (24); cryst (0)
Eukaryota
Protein Name EC#ReferenceOrganismGenBank UniprotPDB/3D
 lytic polysaccharide monooxygenase (CvAA9A) 1.14.99.54
1.14.99.56
pubmed
pubmed
Achaetomiella virescens CBS 547.75 AST24379.1 A0A223GEC9 5NLT[A,B,C,D,E,F]
6YDC[A,B,C,D]
6YDD[A,B]
6YDE[A]
6YDF[A,B]
 lytic cellulose monooxygenase (AfAA9_B; AFUA_4G07850; Afu4g07850) 1.14.99.54
1.14.99.56
pubmed
pubmed
Aspergillus fumigatus Af293 XP_752040.1
EAL90002.1
Q4WP32 5X6A[A,B]
6H1Z[A,B]
6HA5[A,B]
6HAQ[A,B]
 lytic cellulose monooxygenase (AfAA9A; BA78_4697) 1.14.99.54 pubmed
Aspergillus fumigatus var. RP-2014 niveus KEY82006.1    
 lytic cellulose monooxygenase (AfAA9B; BA78_0939) 1.14.99.54
1.14.99.56
pubmed
pubmed
Aspergillus fumigatus var. RP-2014 niveus KEY76044.1    
 lytic cellulose monooxygenase (C4-dehydrogenating) (AN1602.2) 1.14.99.56 pubmed
Aspergillus nidulans FGSC A4 EAA64722.1
ABF50850.1
Q5BCX8  
 lytic cellulose monooxygenase (C1-hydroxylating) (CsLPMO9A; CERSUDRAFT_89533) (probable fragment) 1.14.99.54 doi
Gelatoporia subvermispora QHX39626.1
EMD31056.1
A0A6C0PI79
M2QX02
 
 lytic cellulose monooxygenase (CsLPMO9; CERSUDRAFT_85414) 1.14.99.54 doi
Gelatoporia subvermispora B EMD35432.1 M2RAI8 7EXK[A,B,C,D,E,F]
 lytic cellulose monooxygenase (C1-hydroxylating) (CsLPMO9B; CERSUDRAFT_66688) 1.14.99.54 doi
Gelatoporia subvermispora B EMD35593.1 M2QEW2  
 lytic cellulose polysaccharide monooxygenase 9B (lignocellulose-active) (C1-hydroxylating/C4-dehydrogenating) (Cel61G;GtGH61;GtLPMO9B) (LPMO9B) 1.14.99.54
1.14.99.56
  Gloeophyllum trabeum KUC 8013 AEJ35168.1 F8T947
S7RK00
 
 lytic cellulose monooxygenase (C1-hydroxylating/C4-dehydrogenating) (active on xylogucan and β-glucans) (LPMO9A-2; GtLPMO9A-2) 1.14.99.54
1.14.99.56
pubmed
pubmed
Gloeophyllum trabeum NBRC 6430 BAV57612.1 A0A1C9ZMC5
A0A1C9ZP88
 
  lytic cellulose monooxygenase (C1-hydroxylating) (HETIRDRAFT_166613; HiLPMO9B) (HiLPMO9B) 1.14.99.54   Heterobasidion irregulare TC 32-1 ETW87087.1 W4KMP1 5NNS[A,B]
 lytic cellulose monooxygenase (C1-hydroxylating / C4-dehydrogenating) (LsAA9A) 1.14.99.54
1.14.99.56
pubmed
pubmed
Lentinus similis ALN96977.1 A0A0S2GKZ1 5ACF[A]
5ACG[A]
5ACH[A]
5ACI[A]
5ACJ[A]
5N04[A]
5N05[A]
5NKW[A]
5NLN[A]
5NLO[A]
5NLP[A]
5NLQ[A]
5NLR[A]
5NLS[A]
6YDG[A]
7NIM[A]
7NIN[A]
7PQR[A]
7PTZ[A]
7PXI[A]
7PXJ[A]
7PXK[A]
7PXL[A]
7PXM[A]
7PXN[A]
7PXR[A]
7PXS[A]
7PXT[A]
7PXU[A]
7PXV[A]
7PXW[A]
7PYD[A]
7PYE[A]
7PYF[A]
7PYG[A]
7PYH[A]
7PYI[A]
7PYL[A]
7PYM[A]
7PYN[A]
7PYO[A]
7PYP[A]
7PYQ[A]
7PYU[A]
7PYW[A]
7PYX[A]
7PYY[A]
7PYZ[A]
7PZ0[A]
8S3F[A]
8S3L[A]
 Lytic cellulose monooxygenase (C4-dehydrogenating) (PMO-2;NcPMO-2;NcLPMO9D;GH61-4;NCU01050) (LPMO9D) 1.14.99.56 pubmed
Neurospora crassa OR74A CAD21296.1
EAA32426.1
XP_326543.1
Q1K8B6
Q8WZQ2
4EIR[A,B]
5TKF[A,B,C,D]
5TKG[A,B]
5TKH[A,B]
5TKI[A,B]
7T5C[A,B]
7T5D[A,B]
7T5E[A,B]
  lytic cellulose monooxygenase (C1-hydroxylating) 9F (PMO-03328;LPMO-03328;NcLPMO9F;GH61-6;NCU03328) (LPMO9F) 1.14.99.54   Neurospora crassa OR74A CAD70347.1
EAA26656.1
XP_322586.1
Q1K4Q1
Q873G1
4QI8[A,B]
 lytic cellulose monooxygenase (C1-hydroxylating) (PMO-01867;LPMO-01867;NcLPMO9J;GH61-10;NCU01867;B13N4.070) (LPMO9J) 1.14.99.54   Neurospora crassa OR74A CAE81966.1
EAA36262.1
XP_329057.1
Q6MGH2
Q7SHD9
 
 lytic cellulose polysaccharide monooxygenase (C1-hydroxylating/C4-dehydrogenating) (PMO-3;NcLPMO9M;GH61-13;NcPMO-3;NCU07898) (LPMO9M) 1.14.99.54
1.14.99.56
  Neurospora crassa OR74A EAA33178.1
XP_328604.1
Q7SA19 4EIS[A,B]
  Lytic cellulose monooxygenase (C4-dehydrogenating) (PMO-02916;LPMO-02916;NcLPMO9C;GH61-3;NCU02916) (LPMO9C) 1.14.99.56 pubmed
Neurospora crassa OR74A EAA36362.1
XP_330104.1
Q7SHI8 4D7U[A,B]
4D7V[A,B]
 lytic cellulose monooxygenase (C1-hydroxylating/C4-dehydrogenating) (GH61-2;NCU07760) 1.14.99.54
1.14.99.56
  Neurospora crassa OR74A EAA29018.1
XP_328466.1
Q7S111  
 Lytic cellulose monooxygenase (C4-dehydrogenating) (GH61-1;NcLPMO9A;NCU02240) 1.14.99.56   Neurospora crassa OR74A EAA30263.1
XP_331016.1
Q7S439 5FOH[A]
 lytic cellulose monooxygenase (C1-hydroxylating) (NCU00836) 1.14.99.54   Neurospora crassa OR74A EAA34466.1
XP_325016.1
Q7SCJ5  
 lytic cellulose monooxygenase (C1-hydroxylating) (PMO-08760;LPMO-08760;NcLPMO9E;GH61-5;NCU08760) (LPMO9E) 1.14.99.54   Neurospora crassa OR74A EAA26873.1
XP_330877.1
Q7RWN7  
 Lytic cellulose monooxygenase (C4-dehydrogenating) (PsLPMOB) 1.14.99.56 pubmed
Pestalotiopsis sp. NCi6 ANB32140.1 A0A167KNY8  
 lytic cellulose monooxygenase (C1-hydroxylating/C-4 dehydrogenating) (PsLPMOA) 1.14.99.54
1.14.99.56
  Pestalotiopsis sp. NCi6 ANB32141.1 A0A167KNZ4  
 lytic cellulose monooxygenase (C1-hydroxylating) (Gh61D;PcGH61D;PcLPMO9D) 1.14.99.54   Phanerochaete chrysosporium K-3 BAL43430.1 H1AE14 4B5Q[A,B]
 Lytic cellulose monooxygenase (C1-hydroxylating/C4-dehydrogenating) also active on hemicelluloses (Pa_4_1020;PaLPMO9H) 1.14.99.54
1.14.99.56
  Podospora anserina S mat+ CAP61476.1
CDP27830.1
B2ADG1  
 lytic cellulose monooxygenase (C1-hydroxylating) (Pa_4_7570;PaLPMO9D) 1.14.99.54   Podospora anserina S mat+ CAP66744.1
CDP28479.1
B2ARG6  
 lytic cellulose monooxygenase (C1-hydroxylating) (Pa_1_16300;PaLPMO9E) 1.14.99.54   Podospora anserina S mat+ CAP67740.1
CDP23998.1
B2ATL7  
 Lytic cellulose monooxygenase (C4-dehydrogenating) (Gh61B;Pa_7_3160;PaLPMO9B) (Gh61B) 1.14.99.56 pubmed
Podospora anserina S mat+ CAP68375.1
CDP31846.1
B2AVF1  
 lytic cellulose monooxygenase (C1-hydroxylating) (Pa_6_7780;PaLPMO9F) 1.14.99.54   Podospora anserina S mat+ CAP71839.1
CDP31230.1
B2B403  
 lytic cellulose monooxygenase (C1-hydroxylating/C4-dehydrogenating) (Gh61A;Pa_2_6530;PaLPMO9A) (GH61A) 1.14.99.54
1.14.99.56
pubmed
pubmed
Podospora anserina S mat+ CAP73254.1
CDP25655.1
B2B629  
 lytic cellulose monooxygenase (C1-hydroxylating) (TaAA9; TaAA9A; TaGH61; TaGH61A;T aLPMO9A) 1.14.99.54 pubmed
Thermoascus aurantiacus ACS05720.1
ABW56451.1
G3XAP7 2YET[A,B]
3ZUD[A]
7PU1[A,B]
7PZ3[A]
7PZ4[A]
7PZ5[A]
7PZ6[A]
7PZ7[A]
7PZ8[A]
7Q1K[A]
8B4G[A]
 lytic cellulose monooxygenase (C4-oxidizing) also active on xyloglucan (MtLPMO9J; MYCTH_79765) (MtLPMO9J) 1.14.99.56 pubmed
Thermothelomyces thermophilus ATCC 42464 AEO56016.1 G2Q7A5 9BJQ[A,B]
9BJR[A,B]
9BJS[A,B]
9BJT[A,B]
9D0Z[A,B]
 lytic cellulose monooxygenase (C4-dehydrogenating) (MYCTH_111088) 1.14.99.56 pubmed
Thermothelomyces thermophilus ATCC 42464 AEO56416.1   9FDL[A,B,C]
 lytic cellulose monooxygenase (C1/4-hydroxylating) (StCel61a;MYCTH_46583) (Cel61A) 1.14.99.54
1.14.99.56
pubmed
pubmed
Thermothelomyces thermophilus ATCC 42464 AEO56542.1 G2Q9T3  
 lytic cellulose monooxygenase (C1-hydroxylating) (MtPMO3;MYCTH_92668) 1.14.99.54   Thermothelomyces thermophilus ATCC 42464 AEO56665.1 G2QAB5 5UFV[A,B,C,D,E,F]
 Lytic cellulose monooxygenase (C1-hydroxylating) (MYCTH_112089;MYCTH112089) 1.14.99.54   Thermothelomyces thermophilus ATCC 42464 AEO60271.1 G2QI82  
 lytic cellulose monooxygenase (C1-hydroxylating/C4-dehydrogenating) also active on xylan (MtLPMO9A) (LPMO9A) 1.14.99.54
1.14.99.56
  Thermothelomyces thermophilus C1 AKO82493.1 A0A0H4K9X4  
 multifunctional lytic cellulose, xylan, and xyloglucan lytic monooxygenase (C1-hydroxylating) (TtLPMO9E; TT172_LOCUS3046) 1.14.99.54
1.14.99.-
pubmed
pubmed
Thermothielavioides terrestris LPH172 SPQ20627.1 A0A446BDM9  
 multifunctional lytic cellulose, xylan, and xyloglucan lytic monooxygenase (C4-dehydrogenating) (TtLPMO9A; TT172_LOCUS8210) 1.14.99.56
1.14.99.-
pubmed
pubmed
Thermothielavioides terrestris LPH172 SPQ25791.1 A0A3S4ATQ2  
 lytic cellulose monooxygenase (weak activity) (C1-hydroxylating) (weak activity) (TtLPMO9B; TT172_LOCUS5098) 1.14.99.54 pubmed
Thermothielavioides terrestris LPH172 SPQ22679.1 A0A3S4B613  
 lytic cellulose monooxygenase (C1-hydroxylating) (TtLPMO9G; TT172_LOCUS1635) 1.14.99.54 pubmed
Thermothielavioides terrestris LPH172 SPQ19216.1 A0A446B9M7  
 lytic cellulose monooxygenase (weak activity) (C1-hydroxylating/C4-dehydrogenating) (weak activity) (TtLPMO9T; TT172_LOCUS3045) 1.14.99.54
1.14.99.56
pubmed
pubmed
Thermothielavioides terrestris LPH172 SPQ20626.1 A0A446BDM8  
 lytic cellulose monooxygenase (C1-hydroxylating) (TtLPMO9E; TtAA9E; TtGH61E; THITE_2122979) 1.14.99.54 pubmed
Thermothielavioides terrestris NRRL 8126 XP_003657366.1
ACE10234.1
AEO71030.1
D0VWZ9
G2RGE5
3EII[A,B,C,D]
3EJA[A,B,C,D]
 lytic cellulose monooxygenase (C1-hydroxylating/C4-dehydrogenating) (EGIV; Egl4; EG4; TrLPMO) (formerly endo-β-1,4-glucanase IV) (LPMO9A) 1.14.99.54
1.14.99.56
  Trichoderma reesei RUTC-30 CAA71999.1
EGR52697.1
G0R6T8
O14405
5O2W[A]
5O2X[A]
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