Activities in Family	Modules of approx. 100 residues found in primarily in bacterial pullulanases. The N-terminal module from Thermotoga maritima Pul13 has been shown to bind to the α-glucans amylose, amylopectin, pullulan, and oligosaccharide fragments derived from these polysaccharides (Lammerts van Bueren et al. (2004) Biochemistry 43:15633-42) (PMID: 15581376).
Note	Formerly known as X28 modules.
External resources	CAZypedia;
Statistics	GenBank accession (5166); Uniprot accession (36); PDB accession (20); 3D entries (11); cryst (1)

Bacteria
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D Carbohydrate Ligands Resolution (Å)
multidomain α-amylase (EUR_21100) (cell-wall anchored)		Agathobacter rectalis DSM 17629	CBK91127.1	D6DYI9	6AZ5[A] 2.20
pullulanase I (PulA)		Fervidobacterium pennivorans Ven5	AAD30387.1	Q9XDB5	cryst
C3F39_10140		Klebsiella pneumoniae KPNIH48	AUY19104.1		5YN2[A] 2.30 5YN7[A] 2.59 5YNA[A] 1.96 5YNC[A] 2.32 5YND[A] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 2.23 5YNE[A] 2.20 5YNH[A] 2.05
BB748_23185		Klebsiella pneumoniae Kp_Goe_71070	APM77900.1		6J33[A,B] 1.30 6J35[A,B] 1.84
pullulanase A (PulA; PUL)		Klebsiella pneumoniae UNF5023	CAA36431.1 ACH14985.1	P07206	2FHB[A] α-D-Glcp-(1-4)-α-D-Glcp 1.80 2FHC[A] α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 1.85 2FHF[A] α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 1.65
pullulanase (PulA;HMPREF9690_00740)		Raoultella ornithinolytica 10-5246	EHT13411.1		2YOC[A,B] 2.88
glycogen-degrading enzyme (SpuA;PulA;SP0268)		Streptococcus pneumoniae TIGR4	AAK74446.1 AAT16713.1 ACK14503.1 ACW45222.1 NP_344806.1	A0A0H2UNG0 Q97SQ7	2J44[A] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 2.1 2YA1[A] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-β-D-Glcp 2.25
glycogen-degrading enzyme (SpuA;PulA;SP0268)		Streptococcus pneumoniae TIGR4	AAK74446.1 AAT16713.1 ACK14503.1 ACW45222.1 NP_344806.1	A0A0H2UNG0 Q97SQ7	2J44[A] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 2.1 2YA1[A] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-β-D-Glcp 2.25
PulA		Streptococcus pyogenes M1 GAS SF370	AAK34661.1 NP_269940.1		2J43[A,B] 1.6
PulA		Streptococcus pyogenes M1 GAS SF370	AAK34661.1 NP_269940.1		2J43[A,B] 1.6
pullulanase (PulA;Tm1845) (Pul13A)		Thermotoga maritima MSB8	AAD36907.1 AAM56795.1 ABJ38302.1 ABJ38322.1 ACW20067.1 ACW20078.1 AGL50777.1 AHD18265.1 AKE29594.1 CAA04522.1 CAH68696.1 CAT81460.1 NP_229641.1	G4FGN0 O33840	2J71[A] 1.69 2J72[A,B] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 1.49 2J73[A,B] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-6)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 1.4
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