Activities in Sub Family	β-glucosidase (EC 3.2.1.21);
Mechanism	Retaining e
Clan	GH-A
3D Structure Status	( β / α ) 8 barrel
Catalytic Nucleophile/Base	Glu (experimental)
Catalytic Proton Donor	Glu (experimental)
Note	Once known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resources	CAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)	MEGAZYME; NZYTech; PROZOMIX;
Statistics	GenBank accession (187); Uniprot accession (1);

Bacteria
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D	Subf
GCM10010200_002790		Actinomadura rugatobispora JCM 3366	BFE28028.1			43
HUT06_00695		Actinomadura sp. NAK00032	QKW40094.1			43
SAMN04489716_4612		Actinoplanes derwentensis DSM 43941	SDT57550.1			43
Q0Z83_067810		Actinoplanes sichuanensis 03-723	BEL08590.1			43
L3i22_084230		Actinoplanes sp. L3-i22	BCY13335.1			43
TPMA0078_60650		Actinoplanes sp. TPMA0078	BFD54661.1			43
KCV87_25105		Actinosynnema mirum ATCC 31280	QUF02713.1			43
Amir_6164		Actinosynnema mirum DSM 43827	ACU39971.1			43
APASM_6117		Actinosynnema pretiosum subsp. pretiosum ATCC 31280	AXX33482.1			43
CNX65_30350		Actinosynnema pretiosum X47	ATE57059.1			43
SAMN04489726_6883		Allokutzneria albata DSM 44149	SDN49924.1			43
ISP_002515		Amycolatopsis mediterranei ISP5501	UZF69367.1			43
B737_2348		Amycolatopsis mediterranei RB	AGT83012.1			43
RAM_12075		Amycolatopsis mediterranei S699	AEK40905.1 AFO75883.1			43
AMED_2374		Amycolatopsis mediterranei U32	ADJ44170.1			43
SD37_23615		Amycolatopsis orientalis B-37	ANN18328.1			43
QP939_50585		Amycolatopsis sp. 2-2	WIV56922.1			43
QRY02_01260		Amycolatopsis sp. DG1A-15b	WIX89109.1			43
HUT10_39770		Amycolatopsis sp. Hca4	QKV79233.1			43
F0U60_01135		Archangium minus Cbm 6	WNG52163.1			43
F0U61_01115		Archangium violaceum Cbvi 34	WNG41736.1			43
JQX13_12975		Archangium violaceum SDU34	QRK10897.1			43
JRI60_22000		Archangium violaceum SDU8	QRO01500.1			43
KFF03_16705 (fragment)		Bacterioplanoides sp. SCSIO 12839	UTW48170.1			43
N8J89_28190		Crossiella sp. CA-258035	WHT16995.1			43
HDN1F_08830		gamma proteobacterium HdN1	CBL44466.1			43
KALB_5052		Kutzneria albida DSM 43870	AHH98414.1			43
Q4V64_03070		Kutzneria buriramensis DSM 45791	WKX06526.1			43
JJ691_40870		Kutzneria sp. CA-103260	QUQ66360.1			43
P0136_13075		Lentisphaerota bacterium ZTH	WET06290.1			43
JYG24_09410		Lentisphaerota bacterium zth2	QSH40476.1			43
BBK82_00065		Lentzea guizhouensis DHS C013	ANZ42376.1			43
V1227_15975		Myxococcaceae DG1S-22	WVH84177.1			43
ND450_18885		Lentzea sp. HUAS12	USX56080.1			43
A0128_03105		Leptospira alstonii GWTS #1	AOP35869.1			43
LBF_2411		Leptospira biflexa serovar Patoc strain Patoc 1 (Ames)	ABZ94901.1			43
LEPBI_I2486		Leptospira biflexa serovar Patoc strain Patoc 1 (Paris)	ABZ98573.1			43
EFP84_13910		Leptospira kmetyi LS 001/16	AYV56491.1			43
LPTSP3_g06260		Leptospira kobayashii E30	BDA77696.1			43
HZU44_12355		Micromonospora carbonacea africana	QLK00716.1			43
LV483_02160		Myxococcales bacterium MSr11368	WXA85112.1			43
FNZ07_03730		Paraburkholderia megapolitana LMG 23650	QDQ80348.1			43
V1457_20220		Saccharopolyspora sp. SCSIO 74807	WWR57632.1			43
IOD16_19960		Saccharothrix sp. 6-C	QQQ73565.1			43
LVJ94_02155		Sorangiineae bacterium MSr11367	WXB06066.1			43
LZC94_30015		Sorangiineae bacterium MSr11954	WXB12078.1			43
LZC95_02030		Sorangiineae bacterium MSr12523	WXA95620.1			43
K1J60_02335		Streptomyces akebiae MG28	QYX75507.1			43
HCX60_34365		Streptomyces antibioticus DSM 41481	QIT48001.1			43
QR300_02805		Streptomyces antimycoticus FIM95-F1	WJD95005.1			43
G9272_40365		Streptomyces asoensis H2S5	QJT05810.1			43
O1Q96_29850		Streptomyces aurantiacus Qhu-M197	WAU83530.1			43
SAV_7508		Streptomyces avermitilis MA-4680 = NBRC 14893	BAC75219.1			43
SAVMC3_85070 (fragment)		Streptomyces avermitilis MC3	BBJ55878.1			43
SAVMC3_85060 (fragment)		Streptomyces avermitilis MC3	BBJ55877.1			43
β-glucopyranosidase (SBI_01663)	3.2.1.21	Streptomyces bingchenggensis BCW-1	ADI04784.1	D7CFP8		43
GQF42_39510		Streptomyces broussonetiae T44	QHA08544.1			43
EJC51_41665		Streptomyces brunneus CR22	AZP22036.1			43
IHE65_03325		Streptomyces caniscabiei ID-03-3A	WEO22248.1			43
OG925_06750		Streptomyces canus NBC_01585	WSD84002.1			43
CP983_02855		Streptomyces chartreusis ATCC 14922	QEV72928.1			43
HUT05_04305		Streptomyces chartreusis NA02069	QKZ16657.1			43
STRCI_007432		Streptomyces cinnabarinus DSM 40467	WAZ25904.1			43
CP976_39340		Streptomyces coeruleorubidus ATCC 13740	QEV30738.1			43
PV963_41625		Streptomyces coeruleorubidus DM	WDV56381.1			43
R5U08_39845		Streptomyces coeruleorubidus WT-48	WOT39895.1			43
RFN52_37505		Streptomyces collinus CGMCC 4.1623	WMX68728.1			43
B446_32895		Streptomyces collinus Tu 365	AGS73383.1			43
QU709_00510		Streptomyces coralus SX92	WLW49930.1			43
EJ357_02935		Streptomyces cyaneochromogenes MK-45	AZQ32521.1			43
S1361_35505		Streptomyces cyanogenus S136	QTE02693.1			43
N8I84_37435		Streptomyces cynarae HUAS 13-4	UXY23745.1			43
D9753_02700		Streptomyces dangxiongensis Z022	AYN38038.1			43
BN159_1009		Streptomyces davaonensis JCM 4913	CCK25388.1			43
L3078_41370		Streptomyces deccanensis KCTC 19241	ULR55215.1			43
GFH48_05125		Streptomyces fagopyri QMT-28	QFZ72730.1			43
IM697_20785		Streptomyces ferrugineus CCTCC AA2014009	QOV40623.1			43
CP966_32840		Streptomyces galilaeus ATCC 14969	QEU70510.1			43
GCM10017557_07470		Streptomyces glomeroaurantiacus JCM 4677	BCL25888.1			43
SGFS_101950		Streptomyces graminofaciens A-8890	BBC38901.1			43
AVL59_17735		Streptomyces griseochromogenes ATCC 14511	ANP51215.1			43
DWG14_00902		Streptomyces griseorubiginosus 3E-1	AYC36692.1			43
ELQ87_02750		Streptomyces griseoviridis F1-27	AZS83332.1			43
DDJ31_36610		Streptomyces griseoviridis K61	QCN90965.1			43
CEB94_38125		Streptomyces hawaiiensis ATCC 12236	QCD60988.1			43
V8J11_36520		Streptomyces hygroscopicus subsp. hygroscopicus DSM 41954	WXA10809.1			43
SHXM_09064		Streptomyces hygroscopicus XM201	AQW55601.1			43
RFN58_40220		Streptomyces iakyrus CGMCC 4.1912	WVG06938.1			43
ABB07_02795		Streptomyces incarnatus NRRL 8089	AKJ08988.1			43
SIRAN836		Streptomyces iranensis	CDR02600.1			43
RI060_02315		Streptomyces janthinus JCM 4387	WND16255.1			43
KJK29_01600		Streptomyces koelreuteriae MG62	QWB21367.1			43
JEQ17_06345		Streptomyces liliifuscus ZYC-3	QQM39125.1			43
GJU35_41385		Streptomyces lincolnensis B48	QMV11507.1			43
SLCG_7737		Streptomyces lincolnensis LC-G	AXG58892.1			43
SLINC_7771		Streptomyces lincolnensis NRRL 2936	ANS69995.1			43
SMALA_7592		Streptomyces malaysiensis DSM 4137	ATL87808.1 QDL68827.1			43
R8789_04925		Streptomyces malaysiensis NA3722	WPB88627.1			43
POD33_36910		Streptomyces moderatus DT446	WCH97416.1			43
DVK44_31820		Streptomyces paludis GSSD-12	AXG83034.1			43
OHB35_06375		Streptomyces phaeochromogenes NBC 01752	WSD12889.1			43
OG931_05970		Streptomyces phaeochromogenes NBC_00186	WRZ27326.1			43
F9278_43445		Streptomyces phaeolivaceus GY16	QFR01905.1			43
NFX46_14430		Streptomyces phaeoluteigriseus Qhu-M197	USQ84884.1			43
SPRI_1194		Streptomyces pristinaespiralis HCCB 10218	ALC19500.1			43
H3L99_30995		Streptomyces pristinaespiralis PR11	QMU17479.1			43
BFF78_03590		Streptomyces puniciscabiei TW1S1	AOR36861.1			43
A4E84_37985		Streptomyces qaidamensis S10(2016)	AMW15701.1			43
LUW77_16910		Streptomyces radiopugnans MRC003	URN12701.1			43
LIV37_04095		Streptomyces rapamycinicus NRRL 5491	UTP28600.1			43
M271_03950		Streptomyces rapamycinicus NRRL 5491	AGP52420.1			43
TUE45_00384		Streptomyces reticuli	CUW25674.1			43
IAG44_37005		Streptomyces roseirectus CRXT-G-22	QNP74509.1			43
SCAB_8561		Streptomyces scabiei 87.22	CBG68039.1			43
I1A49_41735		Streptomyces solisilvae HNM0141	QPI60578.1			43
CGL27_46220		Streptomyces sp. 11-1-2	ASQ99437.1			43
P8T65_04350		Streptomyces sp. 11x1	WNZ06904.1			43
SAMN05444521_0940		Streptomyces sp. 3214.6	SHH56234.1			43
IOD14_17460		Streptomyces sp. A2-16	QUC58459.1			43
N7U49_05560		Streptomyces sp. AD2-2	UXX91972.1			43
AS97_19020		Streptomyces sp. AgN23	AJZ86939.2			43
RPQ07_06285		Streptomyces sp. AM2-3-1	WNO71254.1			43
CVT30_03370		Streptomyces sp. AMCC400023	UJV39030.1			43
JIX56_02560		Streptomyces sp. CA-210063	UUU28870.1			43
AS200_05975		Streptomyces sp. CdTB01	ALV38406.1			43
RJD14_32275		Streptomyces sp. CGMCC 4.1456	WNF66965.1			43
VJ737_02510		Streptomyces sp. CGMCC 4.1772	WRT96621.1			43
Q2K21_17970		Streptomyces sp. CGMCC 4.7035	WNB99800.1			43
NNW99_01600		Streptomyces sp. CRCS-T-1	UUA11910.1			43
NNW98_01600		Streptomyces sp. CRLD-Y-1	UUA04285.1			43
V8690_39965		Streptomyces sp. DG1A-41	WWT99868.1			43
JIX55_44540		Streptomyces sp. DSM 40750	UUU26722.1			43
EASAB2608_00731		Streptomyces sp. EAS-AB2608 NBRC 114648	BCM65397.1			43
WJM95_29720		Streptomyces sp. f51	WYH86330.1			43
A6P39_007015		Streptomyces sp. FXJ1.172	WEO93782.1			43
Q7C01_17150		Streptomyces sp. FXY-T5	WMD06026.1			43
C1703_37360		Streptomyces sp. Go-475	AXE90720.1			43
KBP30_05380		Streptomyces sp. Go40/10 Go 40/10	UFR00644.1			43
LUX31_01925		Streptomyces sp. GQFP	UIX28875.1			43
LUV23_41905		Streptomyces sp. HNM0561	UHH22295.1			43
MQE23_40940		Streptomyces sp. HP-A2021	UOB15051.1			43
N8I87_36425		Streptomyces sp. HUAS 15-9	UXY31494.1			43
O1G22_05220		Streptomyces sp. HUAS 2-6	WBO62263.1			43
F3K22_39020		Streptomyces sp. LBUM 1475	QTU66215.1			43
F3K21_39870		Streptomyces sp. LBUM 1480	QTU58171.1			43
F3K20_38915		Streptomyces sp. LBUM 1482	QTU49998.1			43
RKE30_22965		Streptomyces sp. Li-HN-5-11	WNM33043.1			43
MUK60_05455		Streptomyces sp. LRE541	UPZ27309.1			43
CFP59_01079		Streptomyces sp. M56	AUA08991.1			43
R2B38_39850		Streptomyces sp. N50	WOX14620.1			43
QFW82_05340		Streptomyces sp. NA07423	WHX16485.1			43
OG627_03235		Streptomyces sp. NBC_01429	WUX66024.1			43
C6376_25640		Streptomyces sp. P3	AVV47696.1			43
QR97_02615		Streptomyces sp. PBH53	AKN75089.1			43
GPZ77_30280		Streptomyces sp. QHH-9511	QGZ52083.1			43
HYQ63_42740		Streptomyces sp. Rer75	QLH26509.1			43
HEP84_46605		Streptomyces sp. RLB1-33	QIY75413.1			43
FNV59_48690		Streptomyces sp. RLB1-8	QDO64920.1			43
FNV58_49610		Streptomyces sp. RLB1-9	QDO02919.1			43
FNV62_48010		Streptomyces sp. RLB3-17	QDO44789.1			43
FNV60_46435		Streptomyces sp. RLB3-5	QDO54676.1			43
FNV61_49210		Streptomyces sp. RLB3-6	QDN92404.1			43
HEP85_35980		Streptomyces sp. RPA4-2	QIY65991.1			43
FNV63_48215		Streptomyces sp. S1A1-3	QDO34773.1			43
FNV64_49640		Streptomyces sp. S1A1-7	QDN82521.1			43
FNV65_48200		Streptomyces sp. S1A1-8	QDO24653.1			43
FNV66_47965		Streptomyces sp. S1D4-14	QDN72219.1			43
FNV67_49115		Streptomyces sp. S1D4-20	QDN62168.1			43
FNV68_50275		Streptomyces sp. S1D4-23	QDO13227.1			43
N6H00_02255		Streptomyces sp. S465	WAP53867.1			43
SBE_005909		Streptomyces sp. SBE_14.2	WBO82016.1			43
PS467_02810		Streptomyces sp. SCA4-21	WNE94327.1			43
U5640_02825		Streptomyces sp. SS7	WYV84700.1			43
A7X85_06575		Streptomyces sp. ST1015	QZZ25951.1			43
PBV52_01275		Streptomyces sp. T12	WDF35529.1			43
WBG99_01420		Streptomyces sp. TG1A-60	WXC19320.1			43
GLX30_05690		Streptomyces sp. Tu 2975	QIP83639.1			43
PYR72_09275		Streptomyces sp. VNUA24	WEH13885.1			43
SSPO_009300		Streptomyces sporoclivatus NBRC 100767	BBJ38212.1			43
Strvi_4282		Streptomyces violaceusniger Tu 4113	AEM83933.1			43
Sros_5302		Streptosporangium roseum DSM 43021	ACZ88069.1			43
HUF19_05940		Thalassolituus marinus alknpb1M-1	UXD87010.1			43
HUF19_05940		Thalassolituus marinus alknpb1M-1	UXD87010.1			43
SG35_030015		Thalassomonas actiniarum A5K-106	WDE02641.1			43
SG35_030015		Thalassomonas actiniarum A5K-106	WDE02641.1			43
SG34_030670		Thalassomonas viridans XOM25	WDE09132.1			43
SG34_030670		Thalassomonas viridans XOM25	WDE09132.1			43
Tcur_1364		Thermomonospora curvata DSM 43183	ACY96946.1			43
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