Glycoside Hydrolase Family 5 / Subf 16

Activities in Familyendo-β-1,4-glucanase / cellulase (EC 3.2.1.4); endo-β-1,4-xylanase (EC 3.2.1.8); β-glucosidase (EC 3.2.1.21); β-mannosidase (EC 3.2.1.25); β-glucosylceramidase (EC 3.2.1.45); glucan β-1,3-glucosidase (EC 3.2.1.58); licheninase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); glucan endo-1,6-β-glucosidase (EC 3.2.1.75); mannan endo-β-1,4-mannosidase (EC 3.2.1.78); cellulose β-1,4-cellobiosidase (EC 3.2.1.91); steryl β-glucosidase (EC 3.2.1.104); endoglycoceramidase (EC 3.2.1.123); chitosanase (EC 3.2.1.132); β-primeverosidase (EC 3.2.1.149); xyloglucan-specific endo-β-1,4-glucanase (EC 3.2.1.151); endo-β-1,6-galactanase (EC 3.2.1.164); hesperidin 6-O-α-L-rhamnosyl-β-glucosidase (EC 3.2.1.168); β-1,3-mannanase (EC 3.2.1.-); arabinoxylan-specific endo-β-1,4-xylanase (EC 3.2.1.-); mannan transglycosylase (EC 2.4.1.-)
Activities in Sub Family
Mechanism Retaining
ClanGH-A
3D Structure Status( β / α ) 8
Catalytic Nucleophile/BaseGlu (experimental)
Catalytic Proton DonorGlu (experimental)
NoteOnce known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)MEGAZYME; NZYTech; PROZOMIX;
Statistics GenBank accession (28); Uniprot accession (13);
Eukaryota
Protein Name EC#Organism GenBankUniprotPDB/3DSubf
 AN9166.2   Aspergillus nidulans FGSC A4 EAA61999.1 C8VK26
Q5ARB4
  16
 ORF (fragment)   Aspergillus niger AMY96372.1     16
 endo-β-1,6-galactanase (An03g01050) 3.2.1.164 Aspergillus niger CBS 513.88 CAK38078.1 A2QFW8   16
 AO090012000046 (frameshift)   Aspergillus oryzae RIB40 BAE60287.1 Q2UDS8   16
 glycoside hydrolase family 5 protein (Bofut4_p118670.1)   Botrytis cinerea T4 CCD46317.1     16
 unnamed protein product   Fusarium graminearum PH-1 CEF85947.1     16
 β-1,6-galactanase (Gal1;FoGal1) 3.2.1.164 Fusarium oxysporum 12S BAF42338.1 A0ZSY6   16
 ORF   Leptosphaeria maculans JN3 v23.1.3 CBX90804.1 E4ZHH1   16
 ORF   Leptosphaeria maculans JN3 v23.1.3 CBX97708.1 E5A1Z1   16
 ORF (fragment)   Leptosphaeria maculans JN3 v23.1.3 CBY00459.1 E5A9W8   16
 ORF (fragment)   Leptosphaeria maculans JN3 v23.1.3 CBY00460.1 E5A9W9   16
 MG03374.4   Magnaporthe oryzae 70-15 XP_360831.1 G4N8V8   16
 MG05381.4   Magnaporthe oryzae 70-15 XP_360006.1 G4MKY3   16
 MGG_10423   Magnaporthe oryzae 70-15 XP_003719440.1
XP_366204.1
    16
 BN887_04322   Melanopsichium pennsylvanicum 4 CDI55906.1     16
 NCU05882.1   Neurospora crassa OR74A XP_325737.1 Q7S526   16
 Pa_0_1530   Podospora anserina S mat+ CDP29209.1
CAP62136.1
B2AFB8   16
 sscle_01g002110   Sclerotinia sclerotiorum 1980 UF-70 APA05441.1     16
 sr15758   Sporisorium reilianum SRZ2 CBQ69840.1     16
 MYCTH_52514   Thermothelomyces thermophila ATCC 42464 AEO58520.1     16
 THITE_2145591   Thielavia terrestris NRRL 8126 AEO68439.1     16
 UBRO_07788   Ustilago bromivora UB2112 SAM85236.1     16
 ZT1A5_G7597   Zymoseptoria tritici ST99CH_1A5 SMY26155.1     16
 ZT1A5_G5542   Zymoseptoria tritici ST99CH_1A5 SMY24101.1     16
 ZT3D7_G5593   Zymoseptoria tritici ST99CH_3D7 SMQ50440.1     16
 ZT3D7_G7672   Zymoseptoria tritici ST99CH_3D7 SMQ52519.1     16
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