CAZy ~ Carbohydrate-Active enZymes


   
GlycosylTransferase Family 47
  
Known Activitiesheparan -glucuronyltransferase (EC 2.4.1.225); xyloglucan -galactosyltransferase (EC 2.4.1.-); heparan synthase (EC 2.4.1.-); arabinan alph-L-arabinosyltransferase (EC 2.4.2.-).
 
MechanismInverting
 
NoteThe long animal heparan synthases are made of two domains. The N-terminal domain, which adds b-1,4-GlcA residues, belongs to family GT47 while the C-terminal domain, which adds a-1,4-GlcNAc residues, belongs to family GT64. The plant homologs, which have a single domain, display various specificities ranging from xyloglucan b-galactosyltransferase to a pectin b-glucuronyltransferase and arabinan a-L-arabinosyltransferase.
 
Relevant LinksInterPro   PFAM   
 
StatisticsCAZy Entries (174);  GenBank/GenPept (439);  Uniprot (80);
 
TaxonomyEukaryota(174);  
 
 
        
        Eukaryota          
ProteinEC#OrganismGenBank/GenPeptUniProtPDB/3D
Aa_42180 n.d.Arabidopsis arenosaDQ103595
ABA18110.1
   
At2g32740 (GT13) n.d.Arabidopsis thalianaAC003974
NC_003071
AAC04489.1
NP_180833.1
O48843
  
At4g22580 (GT19) n.d.Arabidopsis thalianaBT011242
BT012538
AK226490
AL033545
AL161557
NC_003075
AAR92278.1
AAS99682.1
BAE98632.1
CAA22163.1
CAB79213.1
NP_193989.1
Q9SUW1
  
At5g19670 n.d.Arabidopsis thalianaNC_003076
NP_197468.1
   
At5g62220 (GT18) 2.4.1.-
Arabidopsis thalianaAB019235
NM_125616
BAA97186.1
NP_201028.1
Q9LVB4
  
At3g45400 n.d.Arabidopsis thalianaAL132953
NC_003074
CAB72489.1
NP_190126.1
Q9M3D9
  
At5g03800 n.d.Arabidopsis thalianaAB005235
NC_003076
BAB08605.1
NP_196000.1
Q9FFN2
  
At1g67410 n.d.Arabidopsis thalianaBT028974
NC_003070
ABI93883.1
NP_176908.1
   
At4g38040 n.d.Arabidopsis thalianaAY050989
AY113959
AL035538
AL161592
NM_119965
AAK93666.1
AAM45007.1
CAB37544.1
CAB80469.1
NP_195517.1
Q9SZK4
  
At1g34270/F23M19.7 n.d.Arabidopsis thalianaAC007454
AY054544
AY081684
NM_103149
AAD39607.1
AAK96735.1
AAM10246.1
NP_564443.1
Q9XID1
  
At5g16890 n.d.Arabidopsis thalianaAY056239
AY142556
AL391141
NC_003076
AAL07088.1
AAN13125.1
CAC01702.1
NP_197191.1
Q9LFL2
  
At2g35100 (probable UDP--L-Ara: arabinan -L-arabinosyltransferase;ARAD1) 2.4.2.-
Arabidopsis thalianaAC004667
BT015054
BT015854
AK228986
NC_003071
AAC61825.1
AAT71926.1
AAU94417.1
BAF00874.1
NP_181055.1
O82181
  
At5g44930/K21C13_11 n.d.Arabidopsis thalianaAY048259
BT002634
AB010693
NM_123861
NM_180801
AAK82521.1
AAO11550.1
BAB10875.1
NP_199306.1
NP_851132.1
Q9FLA5
  
At1g68470 (GT17) n.d.Arabidopsis thalianaAC011915
AK118265
NC_003070
AAG52396.1
BAC42883.1
NP_177014.1
Q8GXF5
Q9CA34
  
At3g42180/T27B3.50 n.d.Arabidopsis thalianaBT026120
AL137079
NM_114085
ABG48476.1
CAB68119.1
NP_189804.1
Q9M2N4
  
At5g11610 n.d.Arabidopsis thalianaAK117711
AL163815
NC_003076
BAC42362.1
CAB87721.1
NP_196722.1
Q8GYD4
Q9LYD1
  
At2g32750 (GT12) n.d.Arabidopsis thalianaAC003974
NC_003071
AAC04488.1
NP_180834.1
O48842
  
AT5g61840/mac9_140 (AtGUT2) n.d.Arabidopsis thalianaAY066039
AY087390
AB080693
NM_125578
AB010069
AY054180
AK220976
AAL47406.1
AAM64940.1
BAC20929.1
NP_568941.1
BAB10084.1
AAL06841.1
BAD94554.1
Q9FLS5
Q940Q8
Q8LB70
  
At3g07620/MLP3.7 n.d.Arabidopsis thalianaAC009176
NC_003074
AAF13080.1
NP_187419.1
Q9SSE8
  
At5g25820 n.d.Arabidopsis thalianaAK229111
NC_003076
BAF00988.1
NP_197954.1
   
At4g13990 (GT14) n.d.Arabidopsis thalianaBT006172
Z97335
AL161537
NM_117474
AAP04155.1
CAB10178.1
CAB78441.1
NP_193135.1
Q84R16
O23260
  
At3g03650/T12J13.7 n.d.Arabidopsis thalianaAC009327
BT006471
AK228299
NC_003074
AAF03467.1
AAP21279.1
BAF00242.1
NP_187015.1
Q9SS65
  
At4g16745 n.d.Arabidopsis thalianaAY080672
AY086184
BT002373
AL161544
NC_003075
NM_117776
AAL86348.1
AAM64263.1
AAN86206.1
CAB78716.1
NP_193407.1
NP_567512.1
Q8RXU2
Q8LD60
  
At5g20260 n.d.Arabidopsis thalianaNC_003076
NP_197526.1
   
At1g21480/F24J8_23 n.d.Arabidopsis thalianaAY074870
AY085129
NC_003070
NM_101998
AAL75891.1
AAM61682.1
NP_173568.1
NP_564141.1
Q8S9K6
Q8LEZ9
  
At4g32790/F4D11_10 n.d.Arabidopsis thalianaAK118321
AL022537
AL161582
NC_003075
BAC42936.1
CAA18582.1
CAB79996.1
NP_195005.1
Q8GXB9
O65522
  
At5g41250 (GT20) n.d.Arabidopsis thalianaAB019225
NC_003076
BAB11101.1
NP_198941.1
Q9FHD8
  
At2g20370 (xyloglucan -galactosyltransferase;Mur3;Kam1) 2.4.1.-
Arabidopsis thalianaAC006569
AY195743
BT023734
AK226372
NM_127596
AAD21751.1
AAO39815.1
AAZ23926.1
BAE98519.1
NP_179627.2
Q9SK65
Q7XJ98
  
At1g74680 n.d.Arabidopsis thalianaAY057616
BT002613
NC_003070
NM_106126
AAL14411.1
AAO11529.1
NP_177606.1
NP_565089.1
Q93ZD5
  
At5g22940 n.d.Arabidopsis thalianaNC_003076
AB005243
BT011629
BT014962
NP_197685.1
BAB10615.1
AAS47635.1
AAT47813.1
Q9FFB4
  
At2g28110 (Irx7;Fra8;Fiber8) n.d.Arabidopsis thalianaAC005851
AY081347
BT006256
DQ182567
DQ182568
NC_003071
NM_179782
AAC98455.1
AAL91236.1
AAP13364.1
ABA60868.1
ABA60869.1
NP_180378.1
NP_850113.2
Q9ZUV3
Q8RXC5
  
UDP-Xyl: xylogalacturonan -1,3-xylosyltransferase (Xgd1;At5g33290/F19N2_10) 2.4.2.-
Arabidopsis thalianaAY049234
AY072629
NM_122853
AAK83577.1
AAL62020.1
NP_198314.1
Q94AA9
  
At5g11130 n.d.Arabidopsis thalianaAL360314
NC_003076
CAB96648.1
NP_196674.1
Q9LFP3
  
At2g31990 (GT15) n.d.Arabidopsis thalianaAC006223
BT015812
BT021113
NM_128759
AAD15405.1
AAU94375.1
AAX12883.1
NP_180759.1
Q9SL00
  
At5g25310 n.d.Arabidopsis thalianaNM_122440
NP_197913.1
   
At1g27440 (AtGUT1) n.d.Arabidopsis thalianaAC004557
BT022053
BT025555
NC_003070
AAF99743.1
AAY25465.1
ABF58973.1
NP_174064.1
Q9FZJ1
  
At1g63450 (GT16) n.d.Arabidopsis thalianaNM_105024
NP_176534.1
   
At3g57630 n.d.Arabidopsis thalianaBT011693
BT012627
AK226415
AL049660
NM_115623
AAS49056.1
AAT06446.1
BAE98560.1
CAB41192.1
NP_191322.1
   
At2g29040 (GT11) n.d.Arabidopsis thalianaAC005315
DQ446576
NM_128461
AAC33230.1
ABE65870.1
NP_180468.1
O81072
  
At5g37000 n.d.Arabidopsis thalianaAB024026
NC_003076
BAB08970.1
NP_198516.1
Q9FGV3
  
Unknown (protein for MGC:151773) n.d.Bos taurusBC140568
AAI40569.1
   
-glucuronyltransferase / -N-acetylglucosaminyltransferase (exostosin-2) (Ext2) 2.4.1.225

2.4.1.224
Bos taurusAF089748
BC126570
NM_177496
AAC35386.1
AAI26571.1
NP_803462.1
O77783
  
Unknown (protein for MGC:155111) n.d.Bos taurusBC133540
AAI33541.1
   
ORF (protein for MGC:160120) n.d.Bos taurusBC150061
AAI50062.1
   
Ext2 (K01G5.6/Rib2) n.d.Caenorhabditis elegans Bristol N2U94835
-
AB077851
Z92803
NM_066967
AAC47510.1
AAW12599.1
BAB83878.1
CAB07245.1
NP_499368.1
O01705
O17920
  
Ext-1 / Ri-1 n.d.Caenorhabditis elegans Bristol N2U94834
AB241458
Z73425
NM_069779
AAC47509.1
BAF48989.1
CAB61014.1
NP_502180.1
O01704
Q9U3J6
  
Cr_42180 n.d.Capsella rubellaDQ103594
ABA18105.1
   
-glucuronyltransferase / -N-acetylglucosaminyltransferase (Ext1) 2.4.1.225

2.4.1.224
Cricetulus griseusAF252858
AAF71276.1
Q9JK82
  
CNG01130 n.d.Cryptococcus neoformans var. neoformans JEC21AE017347
AAW44475.1
   
Exo n.d.Cucumis meloAY582736
AAU04753.1
   
EXT1a n.d.Danio rerioBC133117
AY734455
BX005205
AAI33118.1
AAW29033.1
CAM12981.1
   
EXT1b n.d.Danio rerioAY734456
BC125900
AAW29034.1
AAI25901.1
   
Ext2 n.d.Danio rerioAY786508
AAV48783.1
   
EXT1c n.d.Danio rerioAY734457
BC115216
BC155812
AAW29035.1
AAI15217.1
AAI55813.1
   
exostosin-like 3 (Extl3) n.d.Danio rerioAY786436
BC162406.1
AAV51343.1
AAI62406.1
   
CG8433 (Dext2;sotv) n.d.Drosophila melanogasterAF145598
AE003808
AE003808
BT003544
AE003808
NM_166150
NM_176187
NM_176188
AAD38573.1
AAF58055.2
AAF58056.1
AAO39548.1
AAO41379.1
NP_725536.1
NP_788367.1
NP_788368.1
Q9Y169
  
heparan sulfate GlcNAc transferase-I/II (CG15110;Dext3;botv) 2.4.1.225

2.4.1.224
Drosophila melanogasterAF132161
AE003797
AB077850
NM_079066
AAD34749.1
AAF57601.1
BAB89399.1
NP_523790.1
Q9XZ08
  
tout-velu (CG10117;Dext1;ttv) n.d.Drosophila melanogasterAF083889
AE003814
BT021403
AB221351
NM_057883
AAC32397.1
AAF58236.1
AAX33551.1
BAE78509.1
NP_477231.1
Q9V730
O76796
  
RCJMB04_7p22 (fragment) n.d.Gallus gallusAJ851575
CAH65209.1
   
exostosin-like 3 (Extl3;Extr1) 2.4.1.223

2.4.1.224
Homo sapiensU96629
AF001690
AF029231
AF083551
BC006363
BT007353
-
-
-
AB007042
AB011091
AK294475
NM_001440
AAB67602.1
AAC39598.1
AAD01877.1
AAD42041.1
AAH06363.1
AAP36017.1
AAW12588.1
AAW12589.1
AAW12594.1
BAA24080.1
BAA25445.2
BAG57702.1
NP_001431.1
O43909
O00225
  
UDP-GlcUA:-glucuronyltransferase / UDP-GlcNAc: -N-acetylglucosaminyltransferase (exostosin-1;Ext1) 2.4.1.224

2.4.1.225
Homo sapiensS79639
U70539
BC001174
AK313129
NM_000127
AAB62283.1
AAC51154.1
AAH01174.1
BAG35949.1
NP_000118.1
Q16394
Q9BVI9
  
UDP-GlcUA:-glucuronyltransferase / UDP-GlcNAc: -N-acetylglucosaminyltransferase (exostosin-2;Ext2) 2.4.1.224

2.4.1.225
Homo sapiensU62740
U72263
U64511
U67356
BC010058
BC068545
NM_000401
AAB07008.1
AAB62718.1
AAC50764.1
AAC51219.1
AAH10058.1
AAH68545.1
NP_000392.1
Q93063
O15288
  
exostosin-like 1 (Extl1) 2.4.1.224
Homo sapiensU67191
AF083623
AF151391
BC065528
-
AL391650
NM_004455
AAC51141.1
AAD02840.1
AAF73172.1
AAH65528.1
AAW12597.1
CAI17130.1
NP_004446.1
Q92935
  
unnamed protein product (fragment) n.d.Macaca fascicularisAB172049
BAE89111.1
   
MtrDRAFT_AC144540g3v1 n.d.Medicago truncatulaAC144540
AC182813
ABE92985.1
ABP03863.1
   
MtrDRAFT_AC148816g3v1 n.d.Medicago truncatulaAC148816
DQ984942
ABD28410.1
ABK28848.1
   
MtrDRAFT_AC144540g11v1 or MtrDRAFT_AC144657g20v2 n.d.Medicago truncatulaAC144540
AC182813
AC144657
AC144657
ABE92993.1
ABP03873.1
ABE86351.1
ABE86351.2
   
MtrDRAFT_AC148405g21v1 n.d.Medicago truncatulaAC148405
AC148405
ABE81786.1
ABE81786.2
   
MtrDRAFT_AC119408g8v1 n.d.Medicago truncatulaAC119408
ABE86701.1
   
MtrDRAFT_AC148607g4v1 n.d.Medicago truncatulaAC148607
ABE92010.1
   
MtrDRAFT_AC149304g10v1 (fragment) n.d.Medicago truncatulaAC149304
ABE79352.1
   
MtrDRAFT_AC147481g21v1 n.d.Medicago truncatulaAC147481
AC147481
AC147481
ABC75371.1
ABN04853.1
ABN04854.1
   
MtrDRAFT_AC144657g12v2 n.d.Medicago truncatulaAC144657
AC182813
AC144540
ABE86352.1
ABP03872.1
ABE92992.1
   
MtrDRAFT_AC137546g35v1 n.d.Medicago truncatulaAC137546
AC158504
ABE81272.1
ABP03600.1
 &n