Polysaccharide Lyase Family 1

Activities in Familypectate lyase (EC 4.2.2.2); exo-pectate lyase (EC 4.2.2.9); pectin lyase (EC 4.2.2.10).
3D Structure Statusparallel β-helix
External resourcesCAZypedia; HOMSTRAD;
Commercial Enzyme Provider(s)MEGAZYME; PROZOMIX;
Statistics GenBank accession (6001); Uniprot accession (587); PDB accession (42); 3D entries (14); cryst (1)
All (2545) Archaea (17) Bacteria (1934) Eukaryota (552) Viruses (18) unclassified (24) Structure (14 - 1 cryst) Characterized (106)
| 1 | 2 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D Subf
 pectate lyase 4.2.2.2 Bacillus halodurans AAE59748.1
AAS29295.1
    5
 pectate lyase (Pel-4B;BH3819) 4.2.2.2 Bacillus halodurans C-125 BAB07538.1
CAW61441.1
CAW70607.1
CBC41450.1
CBC59731.1
NP_244686.1
Q9K6B0   5
 pectate lyase (Pel-4A;BH0698) 4.2.2.2 Bacillus halodurans C-125 BAB04417.1
CAW40844.1
CAW46002.1
CAW49206.1
CAW50913.1
CAW61440.1
CAW62374.1
CAW70606.1
CAW71262.1
CAW73742.1
CAW79719.1
CAW89137.1
CAW91084.1
CAW91785.1
CAW95934.1
CAW96155.1
CAW97001.1
CAX02078.1
CAX06648.1
CAX07002.1
CAX07505.1
CBC11504.1
CBC98341.1
NP_241564.1
Q9KF01  
 pectate lyase (PelA) 4.2.2.2
4.2.2.9
Bacillus licheniformis 14A CAD56882.1
AAE59746.1
AAN26179.1
AAS29293.1
ACP63662.1
Q8GCB2   5
 BLi04129 or BL00947 (PelII) 4.2.2.2
4.2.2.9
Bacillus licheniformis DSM 13 = ATCC 14580 AAU42942.1
AAU25568.1
CAW40854.1
CAW46012.1
CAW49216.1
CAW50923.1
CAW61466.1
CAW62384.1
CAW70632.1
CAW71272.1
CAW73752.1
CAW79729.1
CAW89163.1
CAW91107.1
CAW91806.1
CAW95944.1
CAW96165.1
CAW97011.1
CAX02088.1
CAX06658.1
CAX07012.1
CAX07515.1
CBC11514.1
CBC98361.1
Q65DC2   5
 pectate lyase (Pel) 4.2.2.2 Bacillus licheniformis RN1 BAG12908.1 Q8GCB2   5
 pectate lyase (Pel-22) 4.2.2.2 Bacillus pumilus BS22 ADK23889.1 D8X181   5
 pectate lyase (PelB) 4.2.2.2 Bacillus pumilus DKS1 ACD11362.1 B2ZFM0   5
 pectate lyase 4.2.2.2 Bacillus sp. AAI12 AAE59747.1
AAS29294.1
    5
 pectate lyase 4.2.2.9 Bacillus sp. I534 AAE59749.1
AAR65348.1
AAS29296.1
    5
 pectate lyase (Pel-103) 4.2.2.2 Bacillus sp. KSM-P103 BAA76885.1
CAW61508.1
CAW70674.1
CBC41578.1
CBC59865.1
Q9WX56   5
 pectate lyase (Pel7;Pel-7) (fragment) 4.2.2.2 Bacillus sp. KSM-P7 BAA76884.1 Q9XE02   5
 pectate lyase (PelA;Bsp165PelA) 4.2.2.2 Bacillus sp. N16-5 ACY38198.1 D0VP31 3VMV[A]
3VMW[A]
5
 pectate lyase (Pel-34K) 4.2.2.2
4.2.2.9
Bacillus sp. P-2850 BAC11008.1 Q8L0S6   5
 pectate lyase (Pel-4B) 4.2.2.2 Bacillus sp. P-4-N BAA96478.1 Q9NDN1   5
 pectate lyase (Pel-4A) 4.2.2.2 Bacillus sp. P-4-N BAA96477.1 Q9NDN2  
 pectate lyase (Pl47) 4.2.2.2 Bacillus sp. TS-47 BAB40336.1
CAW61498.1
CAW70664.1
CBC41559.1
CBC59844.1
Q9AJM4 1VBL[A] 6
 pectate lyase (PelK) 4.2.2.2 Bacillus sp. YA-14 BAA05383.1 Q6LEQ4   6
 polygalacturonate lyase (PelA) 4.2.2.2 Bacillus subtilis 7-3-3 AGN03891.1     6
 pectate lyase (Pel) 4.2.2.2 Bacillus subtilis AC327 BAA22313.1 P39116   6
 pectate lyase (Pel-66) 4.2.2.2 Bacillus subtilis BS66 ADK23888.1 D8X180   6
 pectin lyase (Ppr) 4.2.2.10 Bacillus subtilis IFO 3134 BAA12119.1
CAW40861.1
CAW46019.1
CAW49223.1
CAW50930.1
CAW61472.1
CAW62391.1
CAW70638.1
CAW71279.1
CAW73759.1
CAW79736.1
CAW89178.1
CAW91125.1
CAW91820.1
CAW95951.1
CAW96172.1
CAW97018.1
CAX02095.1
CAX06665.1
CAX07019.1
CAX07522.1
CBC11521.1
CBC98374.1
P94449   8
 pectate lyase (BsPel-PB1) 4.2.2.2 Bacillus subtilis PB1 AOE45992.1     6
 pectate lyase (Pel) 4.2.2.2 Bacillus subtilis RCK AFH66771.1     6
 pectate lyase (Pel) 4.2.2.2 Bacillus subtilis SO113 CAA52866.1 P39116   6
 pectate lyase (Pel;Pel168;BSU07560) 4.2.2.2 Bacillus subtilis subsp. subtilis str. 168 CAB12585.1
AAR45489.1
AIY92046.1
AQR80710.1
AQR84923.1
CAD67509.1
CAD67510.1
CAD67511.1
CAF05441.1
CAW61552.1
CAW70718.1
CBC41660.1
CBC59954.1
NP_388637.1
P39116 1BN8[A]
2BSP[A]
2NZM[A]
2O04[A]
2O0V[A]
2O0W[A]
2O17[A]
2O1D[A]
3KRG[A]
5AMV[A]
6
 pectate lyase D (PelD) 4.2.2.2 Dickeya chrysanthemi B374 CAA35176.1
CAW40922.1
CAW46080.1
CAW49284.1
CAW50991.1
CAW61549.1
CAW62452.1
CAW70715.1
CAW71340.1
CAW73820.1
CAW79797.1
CAW89321.1
CAW91256.1
CAW91914.1
CAW96012.1
CAW96233.1
CAW97079.1
CAX02156.1
CAX06726.1
CAX07080.1
CAX07583.1
CBC11582.1
CBC98484.1
P18209   6
 pectate lyase E (PelE) 4.2.2.2 Dickeya chrysanthemi B374 CAA35175.1
CAW40924.1
CAW46082.1
CAW49286.1
CAW50993.1
CAW61551.1
CAW62454.1
CAW70717.1
CAW71342.1
CAW73822.1
CAW79799.1
CAW89327.1
CAW91260.1
CAW91918.1
CAW96014.1
CAW96235.1
CAW97081.1
CAX02158.1
CAX06728.1
CAX07082.1
CAX07585.1
CBC11584.1
CBC98488.1
P0C1A4
P18210
  6
 pectate lyase (PelA) 4.2.2.2 Dickeya chrysanthemi EC16 AAA24843.1
CAW40921.1
CAW46079.1
CAW49283.1
CAW50990.1
CAW61547.1
CAW62451.1
CAW70713.1
CAW71339.1
CAW73819.1
CAW79796.1
CAW89319.1
CAW91254.1
CAW91911.1
CAW96011.1
CAW96232.1
CAW97078.1
CAX02155.1
CAX06725.1
CAX07079.1
CAX07582.1
CBC11581.1
CBC98482.1
P0C1A2
P29155
1JRG[A,B]
1JTA[A]
1OOC[A,B]
1PE9[A,B]
6
 pectate lyase (PelB;PlB) 4.2.2.2 Dickeya chrysanthemi EC16 AAA24847.1 P04959   3
 pectate lyase (PelC) 4.2.2.2 Dickeya chrysanthemi EC16 AAA24849.1
AAR45490.1
AAW11900.1
P11073 1AIR[A]
1O88[A]
1O8D[A]
1O8E[A]
1O8F[A]
1O8G[A]
1O8H[A]
1O8I[A]
1O8J[A]
1O8K[A]
1O8L[A]
1O8M[A]
1PLU[A]
2EWE[A]
2PEC[A]
3
 pectate lyase (PelE) 4.2.2.2 Dickeya chrysanthemi EC16 AAA24844.1
CAW40923.1
CAW46081.1
CAW49285.1
CAW50992.1
CAW61550.1
CAW62453.1
CAW70716.1
CAW71341.1
CAW73821.1
CAW79798.1
CAW89324.1
CAW91258.1
CAW91916.1
CAW96013.1
CAW96234.1
CAW97080.1
CAX02157.1
CAX06727.1
CAX07081.1
CAX07584.1
CBC11583.1
CBC98486.1
P04960 1PCL[A] 6
 pectate lyase D (PelD;PLd;Dda3937_03372) 4.2.2.2 Dickeya dadantii 3937 ADM99552.1
CAA10570.1
CAW40891.1
CAW46049.1
CAW49253.1
CAW50960.1
CAW61511.1
CAW62421.1
CAW70677.1
CAW71309.1
CAW73789.1
CAW79766.1
CAW89250.1
CAW91190.1
CAW91860.1
CAW95981.1
CAW96202.1
CAW97048.1
CAX02125.1
CAX06695.1
CAX07049.1
CAX07552.1
CBC11551.1
CBC98428.1
E0SAZ3
Q9Z4J4
  6
 pectate lyase Z (PelZ;Dda3937_04191) 4.2.2.2 Dickeya dadantii 3937 ADN00347.1
CAA65785.1
E0SJQ8
P94773
  2
 pectate lyase C (PelC;PLc;Dda3937_04192) 4.2.2.2 Dickeya dadantii 3937 ADN00346.1
CAA10642.1
CAA47822.1
E0SJQ7
Q59420
Q79AJ0
  3
 pectate lyase E (PelE;PLe;Dda3937_03371) 4.2.2.2 Dickeya dadantii 3937 ADM99551.1
AAA24854.1
P0C1A5   6
 pectate lyase A (PelA;Dda3937_03370) 4.2.2.2 Dickeya dadantii 3937 ADM99550.1
AAA24846.1
P0C1A3   6
 pectate lyase B (PelB;Dda3937_04193) 4.2.2.2 Dickeya dadantii 3937 ADN00345.1
CAA47821.1
S25262
E0SJQ6
Q59419
  3
 pectate lyase (Pel-90) 4.2.2.2 Lysinibacillus fusiformis BS90 ADK23887.1 D8X179   6
 pectate lyase (PelB) 4.2.2.2 Paenibacillus amylolyticus 27C64 ADB78775.1 D3JTC2  
 pectate lyase (PelN) 4.2.2.2 Paenibacillus sp. 0602 AGM38211.1    
 ECA4067 (PelA) 4.2.2.2 Pectobacterium atrosepticum SCRI1043 CAG76964.1 Q6CZT4   3
 pectate lyase III (Pel3;PelC) 4.2.2.2 Pectobacterium carotovorum BAA00953.1 P0C1C2
P29171
  3
 pectin lyase (Pnl) 4.2.2.10 Pectobacterium carotovorum AAA24857.1
CAW61559.1
CAW70725.1
CBC41674.1
CBC59969.1
P24112   8
 pectate lyase (Pel-1;Pel1) 4.2.2.2 Pectobacterium carotovorum 71 AAA73933.1 Q47467   3
 pectin lyase (PnlA) (fragment) 4.2.2.10 Pectobacterium carotovorum ECC71 AAA24856.1 Q02726   8
 pectate lyase 1 (Pel1;PelI) 4.2.2.2 Pectobacterium carotovorum ER / IAM1068 / ATROSEPTICA EC / ATROSEPTICA C18 AAC60423.1
AAA24845.1
BAA00155.1
CAA57439.1
P0C1C0
P11430
  3
 pectate lyase 2 (Pel2;PelII) 4.2.2.2 Pectobacterium carotovorum ER / IAM1068 / ATROSEPTICA EC / ATROSEPTICA C18 AAC60422.1
AAA24848.1
CAA57440.1
P0C1C1
P11431
  3
 pectate lyase B (PelB) 4.2.2.2 Pseudoalteromonas haloplanktis 505 AAF86343.1
AAF86343.2
Q9KGY6   5
 pectate lyase A (PelA) 4.2.2.2 Pseudoalteromonas haloplanktis ANT/505 AAF86344.2 Q9KGY5   5
 pectate lyase P (PelP) 4.2.2.2 Pseudomonas amygdali pv. lachrymans AAB17879.1 P72242   6
 pectate lyase (Pel) 4.2.2.2 Pseudomonas fluorescens CY091 AAA93535.1
AAB46399.1
Q59671   6
 pectate lyase (PeL) 4.2.2.2 Pseudomonas marginalis N6301 AAC60448.1
BAA06848.1
CAW61554.1
CAW70720.1
CBC41664.1
CBC59958.1
Q51915   6
 pectin lyase (PnL) 4.2.2.10 Pseudomonas marginalis N6301 BAA06847.1
AAA92512.1
P27027
Q51914
  8
 pectate lyase (Pel;Pstru-4) 4.2.2.2 Pseudomonas viridiflava AAB46398.1
AAC41521.1
ABB55454.1
BAA08077.1
CAW61556.1
CAW70722.1
CBC41668.1
CBC59963.1
Q60140   6
 pectate lyase (Pel) 4.2.2.2 Pseudonocardia sp. AAC38059.1
CAW40858.1
CAW46016.1
CAW49220.1
CAW50927.1
CAW61470.1
CAW62388.1
CAW70636.1
CAW71276.1
CAW73756.1
CAW79733.1
CAW89172.1
CAW91119.1
CAW91814.1
CAW95948.1
CAW96169.1
CAW97015.1
CAX02092.1
CAX06662.1
CAX07016.1
CAX07519.1
CBC11518.1
CBC98368.1
O08454  
 pectate lyase 4.2.2.2 Salipaludibacillus agaradhaerens AAE59745.1
AAS29292.1
    5
 pectate lyase (SCO1880;SCI39.27c) 4.2.2.2 Streptomyces coelicolor A3(2) CAC38815.1
CAW61443.1
CAW70609.1
CBC41453.1
CBC59735.1
NP_626147.1
Q93RU6   6
 pectate lyase (SCO2821;SCBAC17F8.12c) 4.2.2.2 Streptomyces coelicolor A3(2) CAC44284.1
NP_627050.1
Q93J76   5
 pectate lyase (pl-str) 4.2.2.2 Streptomyces sp. S27 ACCC 41168 AER93403.1    
 pectate lyase A (PelA;TM0433) 4.2.2.2 Thermotoga maritima MSB8 AAD35518.1
AGL49355.1
AHD17809.1
AKE28227.1
CAW40843.1
CAW46001.1
CAW49205.1
CAW50912.1
CAW61439.1
CAW62373.1
CAW70605.1
CAW71261.1
CAW73741.1
CAW79718.1
CAW89134.1
CAW91082.1
CAW91782.1
CAW95933.1
CAW96154.1
CAW97000.1
CAX02077.1
CAX06647.1
CAX07001.1
CAX07504.1
CBC11503.1
CBC98340.1
NP_228243.1
Q9WYR4 3ZSC[A]
 pectate lyase (PelB) (alkaline) 4.2.2.2 uncultured bacterium AGS49221.1     6
 XC_1298 4.2.2.2 Xanthomonas campestris pv. campestris str. 8004 AAY48367.1 Q4UX56  
 XC_3590 4.2.2.2 Xanthomonas campestris pv. campestris str. 8004 AAY50632.1 Q4UQP1   6
 pectate lyase (Pel;XCC0645) 4.2.2.2 Xanthomonas campestris pv. campestris str. ATCC 33913 CAW61445.1
AAM39961.1
CBC41457.1
CBC59739.1
NP_636037.1
NP_636037.2
Q8PCR8   6
 pectate lyase II (PelB;XCC2815) 4.2.2.2 Xanthomonas campestris pv. campestris str. ATCC 33913 AAM42087.1
NP_638163.1
Q8P6Z9 2QX3[A,B]
2QXZ[A,B]
2QY1[A,B]
 pectate lyase (PelB;Pl;Pstru-3) 4.2.2.2 Xanthomonas campestris PV. MALVACEARUM STRAIN B414 AAC41522.1 Q56806   6
Eukaryota
Protein Name EC#OrganismGenBank UniprotPDB/3D Subf
 pectate lyase (AN7646.2) 4.2.2.2 Aspergillus nidulans FGSC A4 EAA61832.1
ABF50889.1
CAW61463.1
CAW70629.1
CBC41490.1
CBC59776.1
C8VBZ5
Q5AVN4
  7
 pectin lyase (PL1;AN2569.2) 4.2.2.10 Aspergillus nidulans FGSC A4 EAA64674.1
ABF50856.1
Q5BA61   4
 pectin lyase (AN2331.2) 4.2.2.10 Aspergillus nidulans FGSC A4 ABF50854.1
EAA64442.1
Q5BAU9   4
 pectate lyase (PelA;AN0741.2) 4.2.2.2 Aspergillus nidulans FGSC A4 EAA65383.1
AAA80568.1
ABF50844.1
ABO38859.1
CAW40920.1
CAW46078.1
CAW49282.1
CAW50989.1
CAW61546.1
CAW62450.1
CAW70712.1
CAW71338.1
CAW73818.1
CAW79795.1
CAW89317.1
CAW91252.1
CAW91909.1
CAW96010.1
CAW96231.1
CAW97077.1
CAX02154.1
CAX06724.1
CAX07078.1
CAX07581.1
CBC11580.1
CBC98480.1
Q00645   7
 pectin lyase 2 (PL2) (partial) 4.2.2.10 Aspergillus niger     1IDK[A] 4
 pectin lyase F (PelF) 4.2.2.10 Aspergillus niger CBS 120.49 / N400 CAD34589.1 Q8NJK6   4
 pectin lyase C (PelC) 4.2.2.10 Aspergillus niger CBS 120.49 / N400 AAW03313.1 Q5MBK3   4
 pectin lyase A (PelA) (Pl1A) 4.2.2.10 Aspergillus niger CBS 120.49 / N400 CAA43130.1
CAA39305.1
Q01172 1IDJ[A,B]
1IDK[A]
4
 pectin lyase B (PelB) 4.2.2.10 Aspergillus niger CBS 120.49 / N400 CAA46521.1
CAA01023.1
Q00205 1QCX[A] 4
 pectate lyase A (PlyA) 4.2.2.2 Aspergillus niger CBS 120.49 / N400 CAC33162.1
CAW40883.1
CAW46041.1
CAW49245.1
CAW50952.1
CAW62413.1
CAW71301.1
CAW73781.1
CAW79758.1
CAW89232.1
CAW91172.1
CAW91852.1
CAW95973.1
CAW96194.1
CAW97040.1
CAX02117.1
CAX06687.1
CAX07041.1
CAX07544.1
CBC11543.1
CBC98415.1
Q9C2Z0   7
 An03g00190 (PelB) 4.2.2.10 Aspergillus niger CBS 513.88 CAK37997.1 A2QFN7   4
 pectate lyase I (PlyA;An10g00870) 4.2.2.2 Aspergillus niger CBS 513.88 CAK40523.1 A2QV36   7
 An14g04370 (PelA) 4.2.2.10 Aspergillus niger CBS 513.88 CAK48529.1 A2R3I1   4
 An15g07160 (PelF) 4.2.2.10 Aspergillus niger CBS 513.88 CAK48551.1 A2R6A1   4
 An19g00270 (PelD) 4.2.2.10 Aspergillus niger CBS 513.88 CAK47350.1 A2RBL2   4
 pectin lyase A (PelA) 4.2.2.10 Aspergillus niger EIM-6 AFJ80126.1
AFJ80127.1
    4
 pectin lyase D (PelD) 4.2.2.10 Aspergillus niger N756 AAA32701.1 P22864   4
 pectin lyase A (PNL-ZJ5A) 4.2.2.10 Aspergillus niger ZJ5 ALB05716.1     4
 pectin lyase (pnl-zj5a) 4.2.2.10 Aspergillus niger ZJ5 AIX03726.1     4
 pectin lyase 2 (Pel2) 4.2.2.10 Aspergillus oryzae KBN616 BAB82468.1 Q2TXS4
Q8X1X1
  4
 pectin lyase 1 (Pel1) 4.2.2.10 Aspergillus oryzae KBN616 BAB82467.1 Q2TWM1
Q8X1X2
  4
 pectin lyase 2 (Pel2;AO090010000030) 4.2.2.10 Aspergillus oryzae RIB40 BAE65949.1 Q2TXS4   4
 pectin lyase 1 (Pel1;AO090010000504) 4.2.2.10 Aspergillus oryzae RIB40 BAE66352.1
ABO38857.1
Q2TWM1   4
 pectate lyase B (PelB;Plb) 4.2.2.2 Colletotrichum gloeosporioides AAD09857.1
AFO53515.1
CAW61548.1
CAW70714.1
CBC41652.1
CBC59946.1
O59939   7
 pectin lyase (PnlA) 4.2.2.10 Colletotrichum gloeosporioides AAA21817.1 Q00374   4
 pectate lyase 1 (Pel-1) 4.2.2.2 Colletotrichum gloeosporioides f. sp. malvae AAD43564.1
CAW61510.1
CAW70676.1
CBC41582.1
CBC59869.1
Q9Y892   7
 pectate lyase 2 (Pel-2) 4.2.2.2 Colletotrichum gloeosporioides f. sp. malvae AAD43566.1
CAW61509.1
CAW70675.1
CBC41580.1
CBC59867.1
Q9Y890   7
 pectin lyase 2 (Pnl2;Pnl-2) 4.2.2.10 Colletotrichum gloeosporioides f. sp. malvae AAD43565.1 Q9Y891   4
 pectin lyase (Pnl1;Pnl-1) 4.2.2.10 Colletotrichum gloeosporioides f. sp. malvae AAF22244.1 Q9UUS4   4
 pectate lyase (LLP-52) 4.2.2.2 Lilium longiflorum AAA33398.1
ABM68553.1
CAA78976.1
CAV27791.1
CAW40678.1
CAW40925.1
CAW46083.1
CAW49287.1
CAW50994.1
CAW61388.1
CAW61553.1
CAW62455.1
CAW70719.1
CAW71343.1
CAW73823.1
CAW74515.1
CAW77390.1
CAW79800.1
CAW81674.1
CAW89329.1
CAW91262.1
CAW91920.1
CAW96015.1
CAW96236.1
CAW97082.1
CAX02159.1
CAX06729.1
CAX07083.1
CAX07586.1
CBC11585.1
CBC98490.1
P40973   1
 nodulation pectate lyase (Npl;LjNPL) 4.2.2.2 Lotus japonicus AEX93472.1
AFD54063.1
AFD54065.1
    1
 pectate lyase (PelI;Pl1;MwPl1;Ban17) 4.2.2.2 Musa acuminata AAA Group AAF19195.1
ABG74583.1
CAA63496.1
CAV27777.1
CAW40641.1
CAW40890.1
CAW46048.1
CAW49252.1
CAW50959.1
CAW61354.1
CAW61507.1
CAW62420.1
CAW70673.1
CAW71308.1
CAW73788.1
CAW74481.1
CAW77356.1
CAW79765.1
CAW81640.1
CAW89247.1
CAW91188.1
CAW91859.1
CAW95980.1
CAW96201.1
CAW97047.1
CAX02124.1
CAX06694.1
CAX07048.1
CAX07551.1
CBC11550.1
CBC98426.1
Q43783
Q9SDW4
  1
 pectate lyase 4.2.2.2 Nicotiana tabacum CAA43414.1
CAA47630.1
CAA47631.1
CAW40679.1
CAW40926.1
CAW46084.1
CAW49288.1
CAW50995.1
CAW61389.1
CAW61557.1
CAW62456.1
CAW70723.1
CAW71344.1
CAW73824.1
CAW74516.1
CAW77391.1
CAW79801.1
CAW81675.1
CAW89332.1
CAW91265.1
CAW91922.1
CAW96016.1
CAW96237.1
CAW97083.1
CAX02160.1
CAX06730.1
CAX07084.1
CAX07587.1
CBC11586.1
CBC98492.1
P40972   1
Top


Last update: 2017-10-09 © Copyright 1998-2017
AFMB - CNRS - Université d'Aix-Marseille