Glycoside Hydrolase Family 9

Activities in Familyendoglucanase (EC 3.2.1.4); endo-β-1,3(4)-glucanase / lichenase-laminarinase (EC 3.2.1.6); β-glucosidase (EC 3.2.1.21); lichenase / endo-β-1,3-1,4-glucanase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); cellobiohydrolase (EC 3.2.1.91); xyloglucan-specific endo-β-1,4-glucanase / endo-xyloglucanase (EC 3.2.1.151); exo-β-glucosaminidase (EC 3.2.1.165)
Mechanism Inverting
3D Structure Status( α / α ) 6
Catalytic Nucleophile/BaseAsp (experimental)
Catalytic Proton DonorGlu (experimental)
Noteformerly known as cellulase family E.
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Statistics GenBank accession (3461); Uniprot accession (680); PDB accession (42); 3D entries (16); cryst (0)
All (2525) Archaea (4) Bacteria (1362) Eukaryota (947) unclassified (212) Structure (16) Characterized (170)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 10 |
Eukaryota
Protein Name EC#OrganismGenBank UniprotPDB/3D
 glycoside hydrolase 9 (Eng-5)   Adineta ricciae AKC91349.1    
 endo-1,4-β-glucanase (Egl2)   Allium sativum f. pekinense BAL41418.1    
 endo-β-1,4-glucanase   Amitermes dentatus AGP76397.1    
 endo-β-1,4-glucanase 2   Amitermes foreli AGP76399.1    
 endo-β-1,4-glucanase 1   Amitermes foreli AGP76398.1    
 endo-β-1,4-glucanase (Eg65a;AC-EG65;Eg65-a1) 3.2.1.4 Ampullaria crossean ABD24274.1 A0SGJ5  
 endo-β-1,4-glucanase (Eg65-c4)   Ampullaria crossean ABD24281.1 A0SGK2  
 endo-β-1,4-glucanase (Eg65-c3)   Ampullaria crossean ABD24280.1 A0SGK1  
 endo-β-1,4-glucanase (Eg65-c2)   Ampullaria crossean ABD24279.1 A0SGK0  
 endo-β-1,4-glucanase (Eg65c;Eg65-c1) 3.2.1.4 Ampullaria crossean ABD24278.1 A0SGJ9  
 endo-β-1,4-glucanase (Eg65-b3)   Ampullaria crossean ABD24277.1 A0SGJ8  
 endo-β-1,4-glucanase (Eg65-b2)   Ampullaria crossean ABD24276.1 A0SGJ7  
 endo-β-1,4-glucanase (Eg65b;Eg65-b1) 3.2.1.4 Ampullaria crossean ABD24275.1 A0SGJ6  
 endo-β-1,4-glucanase 2   Anoplotermes schwarzi AGP76401.1    
 endo-β-1,4-glucanase 1   Anoplotermes schwarzi AGP76400.1    
 cellulase   Antipaluria urichi AOV94250.1    
 cellulase   Antipaluria urichi AOV94252.1    
 cellulase   Antipaluria urichi AOV94253.1    
 cellulase   Antipaluria urichi AOV94255.1    
 cellulase   Antipaluria urichi AOV94251.1    
 cellulase   Antipaluria urichi AOV94254.1    
 cellulase   Antipaluria urichi AOV94256.1    
 β-1,4-endoglucanase (Apcel45)   Aplysia kurodai BAX09271.1    
 endo-β-1,4-glucanase 8 / cellulase 1 (Cel1;AtCEL1;EGase;At1g70710) (AtGH9B1) 3.2.1.4 Arabidopsis thaliana AAF09072.1
AAG52329.1
AAK82545.1
AAL67092.1
AAO30718.1
AEE35103.1
CAA67156.1
CAA67157.1
CAK32152.1
CAW91525.1
CAW92415.1
CAW94451.1
CAX00889.1
CAX06019.1
CAX07297.1
CBC93706.1
NP_177228.1
O23696
O23697
Q9CAC1
 
 endo-β-1,4-glucanase 1 / cellulase 2 (Cel2;AtCEL2;EGase;At1g02800) (AtGH9B2) 3.2.1.4 Arabidopsis thaliana AAC16418.1
AAF02887.1
AEE27471.1
BAF00867.1
NP_171779.1
O64949
Q9SRX3
 
 endo-β-1,4-glucanase 14 (At2g44560) (AtGH9B11)   Arabidopsis thaliana AAC27458.1
AAC16102.1
AAM14961.1
AEC10438.1
BAD43652.1
NP_181984.1
Q8S8Q4  
 endo-β-1,4-glucanase 15 (Eg15;At2g44570) (AtGH9B12)   Arabidopsis thaliana AAC27459.1
AEC10439.1
ANM61708.1
BAF01119.1
NP_181985.1
O80497  
 endo-β-1,4-glucanase 18 (At4g09740) (AtGH9B14)   Arabidopsis thaliana CAB39641.1
AEE82789.1
CAB78097.1
NP_192712.1
NP_192712.2
Q9SZ90  
 AT3G42155 (fragment)   Arabidopsis thaliana AEE77726.1    
 endo-1,4-glucanase 3 / cellulase 5 (Cel5;AtCEL5;At1g22880) (AtGH9B4)   Arabidopsis thaliana AAB72171.1
AEE30299.1
AEE30300.1
ANM59006.1
NP_173701.1
O23134
Q2V4L8
 
 endo-β-1,4-glucanase 23 (At4g39000) (AtGH9B17)   Arabidopsis thaliana CAB38820.1
ABE66120.1
ABK28672.1
AEE87006.1
BAC42491.1
CAB80563.1
NP_195611.1
Q8GY58  
 endo-β-1,4-glucanase 24 (Eg24;At4g39010) (AtGH9B18)   Arabidopsis thaliana AAL24307.1
AAN72215.1
AEE87007.1
ANM68016.1
CAB38821.1
CAB80564.1
CAW91518.1
CAW92408.1
CAW94445.1
CAX00882.1
CAX06016.1
CAX07294.1
CBC93700.1
NP_568050.1
Q93YQ7  
 endo-β-1,4-glucanase 22 (At4g38990) (AtGH9B16)   Arabidopsis thaliana CAB38819.1
AEE87005.1
ANM67304.1
CAB80562.1
CAW91527.1
CAW92417.1
CAW94453.1
CAX00891.1
CAX06020.1
CBC93708.1
NP_195610.1
Q9SVJ4  
 endo-β-1,4-glucanase 19 (Eg19;At4g11050) (AtGH9C3)   Arabidopsis thaliana AAM91619.1
AAC35539.1
AEE82966.1
ANM66179.1
ANM66180.1
CAB43040.1
CAB81206.1
NP_192843.2
Q8L7I0  
 endo-β-1,4-glucanase 10 (At2g44540) (AtGH9B9)   Arabidopsis thaliana AAC16100.1
AAC27456.1
AAM14965.1
AEC10436.1
NP_181982.1
O64889  
 endo-β-1,4-glucanase 13 (At2g44550) (AtGH9B10)   Arabidopsis thaliana AAC16101.1
AAC27457.1
AAM14964.1
AEC10437.1
NP_181983.1
O64890  
 endo-β-1,4-glucanase 7 (Kor2;At1g65610/F5I14.14) (AtGH9A2)   Arabidopsis thaliana AAB60922.1
AEE34400.1
BAF01264.1
NP_176738.1
O04478  
 endo-β-1,4-glucanase 20 (At4g23560/F9D16.30) (AtGH9B15)   Arabidopsis thaliana CAA23022.1
AEE84775.1
CAB79311.1
NP_194087.1
Q9SUS0  
 endo-β-1,4-glucanase 11 (Eg11;At2g32990) (AtGH9B8)   Arabidopsis thaliana AAB91971.1
AAL32517.1
AAP68324.1
AEC08771.1
NP_180858.1
O48766  
 endo-β-1,4-glucanase 21 (Kor3;At4g24260) (AtGH9A3)   Arabidopsis thaliana CAB45061.1
AEE84879.1
ANM67982.1
ANM67983.1
CAB79336.1
NP_194157.1
Q9STW8  
 endo-β-1,4-glucanase 4 (Eg4;At1g23210) (AtGH9B6)   Arabidopsis thaliana AAC00616.1
AAF86995.1
AAY78616.1
ABH04486.1
AEE30357.1
NP_173735.1
O49296  
 endo-β-1,4-glucanase 17 (Eg17;At4g02290) (AtGH9B13)   Arabidopsis thaliana AAC28173.1
AAL85001.1
AAM26639.1
AEE82151.1
ANM67028.1
CAB80722.1
NP_192138.1
O81416  
 endo-β-1,4-glucanase 9 / cellulase 3 (Cel3;AtCEL3;At1g71380) (AtGH9B3)   Arabidopsis thaliana AAG51817.1
AAA90944.1
AAF23348.1
AAM63253.1
ABH04566.1
AEE35195.1
NP_177294.1
Q0V7W1
Q38817
Q8LDE8
Q9C9H5
 
 endo-β-1,4-glucanase 16 (At3g43860) (AtGH9A4)   Arabidopsis thaliana AAL59921.1
AEE77836.1
ANM65868.1
BAF01489.1
CAB83158.1
NP_189972.2
Q8VYG3
Q9LZG2
 
 endo-β-1,4-glucanase 5 (Eg5;AtCel6;At1g48930) (AtGH9C1)   Arabidopsis thaliana AAF69707.1
AAG29742.1
AAO22749.1
AAO64058.1
AEE32368.1
NP_175323.1
Q9FVQ2
Q9M995
 
 endo-β-1,2-glucanase 2 (At1g19940) (AtGH9B5)   Arabidopsis thaliana AAG12562.1
AAF79918.1
AAK82507.1
AAM47371.1
AEE29915.1
ANM59312.1
ANM59313.1
NP_173423.1
Q9FXI9
Q9LNS3
 
 endo-β-1,4-glucanase 25 (Kor1;Tsd1;KORRIGAN1;AtKOR;At5g49720) (AtGH9A1) 3.2.1.4 Arabidopsis thaliana AAC83240.1
AAB60304.1
AAC33467.1
AAC35344.1
AAK59818.1
AAM63370.1
AAN72232.1
AED95850.1
BAA98160.1
BAD95336.1
BAH57006.1
NP_199783.1
Q38890
Q8H0S4
Q8LD74
Q94C24
 
 endo-β-1,4-glucanase 10 (Eg10;At1g75680) (AtGH9B7)   Arabidopsis thaliana AAF87112.1
AAK64042.1
AAM63477.1
AAN12892.1
AEE35745.1
NP_177697.1
Q8LCP6  
 endo-β-1,4-glucanase 6 (Eg6;At1g64390) (AtGH9C2/AtGH9A10)   Arabidopsis thaliana AAG51703.1
AAG21508.1
AAK50080.1
AAN28884.1
AAN31840.1
AEE34236.1
NP_176621.1
Q42059
Q8H160
 
 glycoside hydrolase family 9 (Gh9-3)   Aretaon asperrimus AMH40358.1    
 glycoside hydrolase family 9 (Gh9-2)   Aretaon asperrimus AMH40357.1    
 glycoside hydrolase family 9 (Gh9-6)   Aretaon asperrimus AMH40361.1    
 glycoside hydrolase family 9 (Gh9-5)   Aretaon asperrimus AMH40360.1    
 glycoside hydrolase family 9 (Gh9-1)   Aretaon asperrimus AMH40356.1    
 glycoside hydrolase family 9 (Gh9-4)   Aretaon asperrimus AMH40359.1    
 ORF (fragment)   Arundinaria densifolia AIM44154.1    
 ORF (fragment)   Arundinaria densifolia AIM44185.1    
 ORF (fragment)   Arundinaria gigantea AIM44171.1    
 ORF (fragment)   Arundinaria gigantea AIM44202.1    
 ORF (fragment)   Arundinaria tecta AIM44203.1    
 ORF (fragment)   Arundinaria tecta AIM44166.1    
 ORF (fragment)   Aulonemia queko AIM44147.1    
 ORF (fragment)   Aulonemia queko AIM44131.1    
 EG (fragment)   Australoplax tridentata AIT38262.1    
 endoglucanase-like (fragment)   Austrothelphusa transversa ABA87134.1 Q38Q22  
 EG (fragment)   Austruca perplexa AIT38263.1    
 endo-β-1,4-glucanase (Cel1) (fragment)   Avena strigosa ADQ73896.1 E5G610  
 ORF (fragment)   Bambusa vulgaris AIM44123.1    
 ORF (fragment)   Bambusa vulgaris AIM44140.1    
 ORF (fragment)   Bambusa vulgaris AIM44139.1    
 ORF (fragment)   Bambusa vulgaris AIM44138.1    
 endo-1,4-β-glucanase (Bc-eg70A)   Bellamya sp. UM-2014 BAS18736.1    
 endo-1,4-β-glucanase (Bc-eg70B)   Bellamya sp. UM-2014 BAU45387.1    
 endo-β-1,4-glucanase (Bc-eg70C)   Bellamya sp. UM-2014 BAU45388.1    
 ORF (fragment)   Bergbambos tessellata AIM44153.1    
 ORF (fragment)   Bergbambos tessellata AIM44187.1    
 endo-1,4-β-glucanase 1 (Kor1)   Betula luminifera AGV22123.1    
 endo-β-1,4-glucanase 2 (Kor2)   Betula luminifera AGV22124.1    
 endo-β-1,4-glucanase 3 (Kor3)   Betula luminifera AGV22125.1    
 β-1,4-endoglucanase (GH9) (fragment)   Biomphalaria glabrata NHM1742 AAT76428.1 Q5ISJ1  
 ORF (fragment)   Brachyelytrum erectum AIM44114.1    
 cellulase (Cel16)   Brassica napus CAB51903.1 Q9STD9  
 unnamed protein product   Brassica napus CBD34635.1    
 ORF (fragment)   Buergersiochloa bambusoides AIM44116.1    
 ORF (fragment)   Buergersiochloa bambusoides AIM44115.1    
 ORF   Camellia sinensis AHB32109.1    
 6J23.18   Capsella rubella ACC91269.1 B2WS97  
 endo-1,4-glucanase (Ccel1;Ccx1)   Capsicum annuum CAA65826.1
CAA58684.1
Q43749
Q96545
 
 endo-β-1,4-glucanase (Ccel2;Eg2;Ccx2)   Capsicum annuum CAA65828.1
CAA58685.1
CAB59900.1
Q43750
Q96546
Q9SML6
 
 endo-β-1,4-glucanase (Ccel3;Ccx3)   Capsicum annuum CAA65827.1
CAA58686.1
CAW91545.1
CAW92434.1
CAW94472.1
CAX00909.1
CAX06028.1
CAX07306.1
CBC93723.1
Q43751
Q96547
 
 endo-β-1,4-glucanase C (Cel1)   Capsicum annuum CAA60737.1 Q42660  
 Cel1 (fragment)   Capsicum chinense BAG30912.1 B2NIX4  
 Cel2 (fragment)   Capsicum chinense BAG30913.1 B2NIX5  
 ORF (fragment)   Cenchrus compressus AFN16446.1    
 membrane-bound endo-1,4-β-D-glucanase, partial (Cokor1) (fragment)   Chamaecyparis obtusa BAN10311.1    
 endoglucanase (Cel1;EG) 3.2.1.4 Cherax quadricarinatus AAD38027.1
AAO61672.2
Q86GS4
Q9Y0W2
 
 ORF (fragment)   Chimonocalamus montanus AIM44155.1    
 ORF (fragment)   Chimonocalamus montanus AIM44186.1    
 ORF (fragment)   Chusquea bambusoides AIM44151.1    
 ORF (fragment)   Chusquea bambusoides AIM44134.1    
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