Glycoside Hydrolase Family 9

Activities in Familyendoglucanase (EC 3.2.1.4); endo-β-1,3(4)-glucanase / lichenase-laminarinase (EC 3.2.1.6); β-glucosidase (EC 3.2.1.21); lichenase / endo-β-1,3-1,4-glucanase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); cellobiohydrolase (EC 3.2.1.91); xyloglucan-specific endo-β-1,4-glucanase / endo-xyloglucanase (EC 3.2.1.151); exo-β-glucosaminidase (EC 3.2.1.165)
Mechanism Inverting
3D Structure Status( α / α ) 6
Catalytic Nucleophile/BaseAsp (experimental)
Catalytic Proton DonorGlu (experimental)
Noteformerly known as cellulase family E.
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Statistics GenBank accession (3581); Uniprot accession (679); PDB accession (42); 3D entries (16); cryst (0)
All (2645) Archaea (4) Bacteria (1479) Eukaryota (950) unclassified (212) Structure (16) Characterized (170)
| 1 | ... | 5 | 6 | 7 | 8 | 9 | 10 | 11 | 12 | 13 | ... | 15 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D
 β-glucan hydrolase (BgH)   Paenibacillus sp. WRL301 BAL46914.1    
 endo-β-1,4-glucanase (Cel9K)   Paenibacillus sp. X4 AUZ62366.1    
 SY83_01955   Paenibacillus swuensis DY6 ANE45298.1    
 BS614_09745   Paenibacillus xylanexedens PAMC 22703 APO44258.1    
 Palpr_0628   Paludibacter propionicigenes WB4 ADQ78784.1 E4T240  
 Palpr_0669   Paludibacter propionicigenes WB4 ADQ78825.1 E4T281  
 Palpr_1774   Paludibacter propionicigenes WB4 ADQ79913.1 E4T5B9  
 Palpr_2972   Paludibacter propionicigenes WB4 ADQ81101.1 E4T0N7  
 BSF38_00374   Paludisphaera borealis PX4 APW58961.1    
 SAMN04489860_1353   Paraoerskovia marina DSM 22126 SDS34914.1    
 S100892_01669 (fragment)   Pediococcus pentosaceus SRCM100892 ARW20213.1    
 S100892_01670 (fragment)   Pediococcus pentosaceus SRCM100892 ARW20214.1    
 AY601_2786   Pedobacter cryoconitis PAMC 27485 AMP99669.1    
 Phep_4258   Pedobacter heparinus DSM 2366 ACU06449.1 C6XXE3  
 Phep_0675   Pedobacter heparinus DSM 2366 ACU02897.1 C6Y1C1  
 BFS30_05470   Pedobacter steynii DX4 AOM76655.1    
 JBW_03171   Pelosinus fermentans JBW45 AJQ28512.1    
 UFO1_3676   Pelosinus sp. UFO1 AIF53219.1    
 CAY62_01660   Photobacterium damselae subsp. damselae KC-Na-1 ARR48395.1    
 PDPUS_1_00526   Photobacterium damselae subsp. piscicida 91-197 BAX51901.1    
 PDP_0018   Photobacterium damselae subsp. piscicida NCIMB 2058 AEW28957.1    
 H744_2c3274   Photobacterium gaetbulicola Gung47 AJR09916.1    
 H744_2c1634   Photobacterium gaetbulicola Gung47 AJR08307.1    
 exo-β-glucosaminidase (GlcNase;PpGlcNase;PBPRA0520) 3.2.1.165 Photobacterium profundum SS9 CAG18943.1   5DGQ[A,B]
5DGR[A,B]
 PSMK_10370   Phycisphaera mikurensis NBRC 102666 BAM03196.1    
 VT85_24155 (CelD)   Planctomyces sp. SH-PL62 AMV40545.1    
 CIK06_08520   Plantactinospora sp. KBS50 ASW57530.1    
 E05_45220 (fragment)   Plautia stali symbiont BAN99288.1    
 E05_45230 (fragment)   Plautia stali symbiont BAN99289.1    
 SAMEA2665130_00497   Plesiomonas shigelloides NCTC10360 SBT60068.1    
 LPB03_05815   Polaribacter vadi LPB0003 AOW17008.1    
 PKOR_09320   Pontibacter korlensis X14-1T AKD03288.1    
 PRU_2222   Prevotella ruminicola 23 ADE81014.1 D5EV45  
 ORF   Prevotella sp. Sc00028 AGH14023.1    
 PSM36_2018   Proteiniphilum saccharofermentans SCD20825.1    
 PSM36_2018   Proteiniphilum saccharofermentans SCD20825.1    
 PESP_a2168   Pseudoalteromonas espejiana ATCC 29659 ASM50178.1    
 CPA52_09205   Pseudoalteromonas issachenkonii ECSMB14103 ATG58409.1    
 ATS72_002890   Pseudoalteromonas sp. 13-15 AUL72605.1    
 glycoside hydrolase GH5   Pseudoalteromonas sp. An33 CUV01081.1    
 cellulase (Cel-JTS)   Pseudomonas aeruginosa ACX31080.1 D0ESZ3  
 endoglucanase (fragment) (Cel9A) 3.2.1.4 Pseudomonas sp. SK38 AAG49558.1 Q9APG3  
 endo-β-1,4-glucanase (CelA) 3.2.1.4 Pseudomonas sp. YD-15 AAD01959.1 Q9Z3X7  
 Psesu_1198   Pseudoxanthomonas suwonensis 11-1 ADV27046.1    
 Psesu_1546   Pseudoxanthomonas suwonensis 11-1 ADV27393.1    
 Psesu_2689   Pseudoxanthomonas suwonensis 11-1 ADV28517.1    
 WQ53_10030   Pseudoxanthomonas suwonensis J1 AKC87029.1    
 WQ53_13890   Pseudoxanthomonas suwonensis J1 AKC87683.1    
 TE10_14310   Raoultella ornithinolytica S12 AJF75294.1    
 AWN76_010685   Rhodothermaceae bacterium RA ARA95103.1    
 Rmar_0076   Rhodothermus marinus DSM 4252 ACY46985.1 D0MK72  
 Rmar_0525   Rhodothermus marinus DSM 4252 ACY47427.1 D0MEX7  
 Rhom172_0073   Rhodothermus marinus SG0.5JP17-172 AEN72024.1    
 Riv7116_5208   Rivularia sp. PCC 7116 AFY57603.1    
 RD2015_2615   Roseateles depolymerans KCTC 42856 ALV07080.1    
 RD2015_3374   Roseateles depolymerans KCTC 42856 ALV07831.1    
 TH63_14730   Rufibacter sp. DG31D AKQ46601.1    
 TH63_14845   Rufibacter sp. DG31D AKQ46618.1    
 DC20_20840   Rufibacter tibetensis strain 1351 ALJ00989.1    
 DC20_05220   Rufibacter tibetensis strain 1351 ALI98485.1    
 DC20_06385   Rufibacter tibetensis strain 1351 ALJ01265.1    
 AD2_01829   Ruminiclostridium thermocellum AD2 ALX08819.1    
 AD2_01980   Ruminiclostridium thermocellum AD2 ALX08970.1    
 AD2_03060   Ruminiclostridium thermocellum AD2 ALX10036.1    
 AD2_00353   Ruminiclostridium thermocellum AD2 ALX07361.1    
 AD2_01828   Ruminiclostridium thermocellum AD2 ALX08818.1    
 AD2_01489   Ruminiclostridium thermocellum AD2 ALX08482.1    
 AD2_00354   Ruminiclostridium thermocellum AD2 ALX07362.1    
 AD2_01409   Ruminiclostridium thermocellum AD2 ALX08402.1    
 AD2_00404   Ruminiclostridium thermocellum AD2 ALX07412.1    
 AD2_01677   Ruminiclostridium thermocellum AD2 ALX08667.1    
 AD2_01614   Ruminiclostridium thermocellum AD2 ALX08607.1    
 AD2_02219   Ruminiclostridium thermocellum AD2 ALX09207.1    
 AD2_01712   Ruminiclostridium thermocellum AD2 ALX08702.1    
 AD2_02216   Ruminiclostridium thermocellum AD2 ALX09204.1    
 AD2_01615   Ruminiclostridium thermocellum AD2 ALX08608.1    
 AD2_01808   Ruminiclostridium thermocellum AD2 ALX08798.1    
 endo-β-1,4-glucanase I (CelI;Cthe_0040) (Cel9I) 3.2.1.4 Ruminiclostridium thermocellum ATCC 27405 ABN51281.1 Q02934  
 Cthe_0043 (CelN) (Cel9N)   Ruminiclostridium thermocellum ATCC 27405 ABN51284.1 A3DBF5  
 Cthe_0274 (CelP)   Ruminiclostridium thermocellum ATCC 27405 ABN51513.1 A3DC34  
 cellulase K (CelK;Cthe_0412) (Cel9K) 3.2.1.4 Ruminiclostridium thermocellum ATCC 27405 ABN51650.1 A3DCH1  
 cellobiohydrolase (CbhA;Cthe_0413) (Cbh9A) 3.2.1.91 Ruminiclostridium thermocellum ATCC 27405 ABN51651.1
AAR87745.1
A3DCH2
Q6RSN8
1RQ5[A]
1UT9[A]
 Cthe_0433   Ruminiclostridium thermocellum ATCC 27405 ABN51671.1 A3DCJ2  
 endo-β-1,4-glucanase F (CelF;Cthe_0543) (Cel9B) 3.2.1.4 Ruminiclostridium thermocellum ATCC 27405 ABN51779.1 P26224  
 endo-β-1,4-glucanase R / cellotetraohydrolase (CelR;Cthe_0578) (Cel9R) 3.2.1.4 Ruminiclostridium thermocellum ATCC 27405 ABN51814.1 A3DCY5  
 broad substrate-specificity xyloglucanase J / processive endoglucanase (CelJ;Cthe_0624) (Cel9D/Cel9J+Cel44A) 3.2.1.151 Ruminiclostridium thermocellum ATCC 27405 ABN51859.1 A3DD30  
 endo-β-1,4-glucanase Q (CelQ;Cthe_0625) (Cel9I) 3.2.1.4 Ruminiclostridium thermocellum ATCC 27405 ABN51860.1 A3DD31  
 Cthe_0745 (CelW)   Ruminiclostridium thermocellum ATCC 27405 ABN51980.1 A3DDF1  
 endo-β-1,4-glucanase (CelD;Cthe_0825) (Cel9A) 3.2.1.4 Ruminiclostridium thermocellum ATCC 27405 ABN52060.1 A3DDN1  
 Cthe_2360 (CelU)   Ruminiclostridium thermocellum ATCC 27405 ABN53562.1 A3DHY3  
 Cthe_2760 (CelV) (Cel9V)   Ruminiclostridium thermocellum ATCC 27405 ABN53959.1 A3DJ30  
 Cthe_2761 (LecA)   Ruminiclostridium thermocellum ATCC 27405 ABN53960.1 A3DJ31  
 endo-β-1,4-glucanase T (CelT;Cthe_2812) (Cel9T) 3.2.1.4 Ruminiclostridium thermocellum ATCC 27405 ABN54011.1 A3DJ82 2YIK[A]
 Clo1313_1694   Ruminiclostridium thermocellum DSM 1313 ADU74751.1    
 Clo1313_0350   Ruminiclostridium thermocellum DSM 1313 ADU73440.1    
 Clo1313_1477   Ruminiclostridium thermocellum DSM 1313 ADU74539.1    
 Clo1313_1396   Ruminiclostridium thermocellum DSM 1313 ADU74459.1    
 Clo1313_1809   Ruminiclostridium thermocellum DSM 1313 ADU74865.1    
 Clo1313_1659   Ruminiclostridium thermocellum DSM 1313 ADU74716.1    
 Clo1313_1603   Ruminiclostridium thermocellum DSM 1313 ADU74664.1    
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