Glycoside Hydrolase Family 9

Activities in Familyendoglucanase (EC 3.2.1.4); endo-β-1,3(4)-glucanase / lichenase-laminarinase (EC 3.2.1.6); β-glucosidase (EC 3.2.1.21); lichenase / endo-β-1,3-1,4-glucanase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); cellobiohydrolase (EC 3.2.1.91); xyloglucan-specific endo-β-1,4-glucanase / endo-xyloglucanase (EC 3.2.1.151); exo-β-glucosaminidase (EC 3.2.1.165)
Mechanism Inverting
3D Structure Status( α / α ) 6
Catalytic Nucleophile/BaseAsp (experimental)
Catalytic Proton DonorGlu (experimental)
Noteformerly known as cellulase family E.
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Statistics GenBank accession (3491); Uniprot accession (679); PDB accession (42); 3D entries (16); cryst (0)
All (2555) Archaea (4) Bacteria (1392) Eukaryota (947) unclassified (212) Structure (16) Characterized (170)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | ... | 14 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D
 cellulase P (P90;Ccel_0753) (Cel9P) 3.2.1.4 [Clostridium] cellulolyticum H10 ACL75131.1
ABG76972.1
Q0PRM9  
 cellulase M (CelM;Ccel_0737) (Cel9M) 3.2.1.4 [Clostridium] cellulolyticum H10 ACL75116.1
AAG45160.1
B8I7V9
Q9EYQ2
1IA6[A]
1IA7[A]
 cellulase J (CelJ;Ccel_0735) (Cel9J) 3.2.1.4 [Clostridium] cellulolyticum H10 ACL75114.1
AAG45158.1
B8I7V7
Q9EYQ4
 
 cellulase H (CelH;Ccel_0734) (Cel9H) 3.2.1.4 [Clostridium] cellulolyticum H10 ACL75113.1
AAG45157.1
B8I7V6
Q9EYQ5
 
 CCDG5_1676   [Clostridium] cellulosi CDZ24784.1    
 CCDG5_1679 (CelZ)   [Clostridium] cellulosi CDZ24787.1    
 endoglucanase (Eng9D)   Clostridium cellulovorans BAV13039.1    
 endoglucanase (Eng9E)   Clostridium cellulovorans BAV13140.1    
 endoglucanase (Eng9C)   Clostridium cellulovorans BAV13038.1    
 endoglucanase (Eng9B)   Clostridium cellulovorans BAV13037.1    
 endoglucanase O (EngO) 3.2.1.4 Clostridium cellulovorans AAT66046.1 Q6DTY2  
 Clocel_1477   Clostridium cellulovorans 743B ADL51228.1 D9SW84  
 Clocel_1476   Clostridium cellulovorans 743B ADL51227.1 D9SW83  
 Clocel_2576   Clostridium cellulovorans 743B ADL52288.1 D9SQT2  
 endo-β-1,4-glucanase (EngZ;NC1;Clocel_2741) 3.2.1.4 Clostridium cellulovorans 743B ADL52438.1
YP_003844202.1
D9SRK9  
 Clocel_0930   Clostridium cellulovorans 743B ADL50698.1 D9ST82  
 Clocel_1420   Clostridium cellulovorans 743B ADL51173.1 D9SW29  
 Clocel_1478   Clostridium cellulovorans 743B ADL51229.1 D9SW85  
 Clocel_1624   Clostridium cellulovorans 743B ADL51370.1 D9SX09  
 Clocel_2816   Clostridium cellulovorans 743B ADL52513.1 D9SS66  
 Clocel_2821 (EngK)   Clostridium cellulovorans 743B ADL52517.1 D9SS70  
 Clocel_2822 (EngH)   Clostridium cellulovorans 743B ADL52518.1 D9SS71  
 endo-β-1,4-glucanase K (EngK) 3.2.1.4 Clostridium cellulovorans 743B AAF06107.1 Q9RGE8  
 endo-β-1,4-glucanase L (EngL;Clocel_2819) 3.2.1.4 Clostridium cellulovorans 743B AAF06109.1
ADL52515.1
C5RQ37
D9SS68
Q9RGE6
 
 endo-β-1,4-glucanase M (EngM) 3.2.1.4 Clostridium cellulovorans 743B AAF06111.2 Q9LAJ2  
 endo-β-1,4-glucanase Y (EngY) 3.2.1.4 Clostridium cellulovorans ATCC 35269 AAG59608.1 Q9AQF4  
 endo-β-1,4-glucanase H (EngH) 3.2.1.4 Clostridium cellulovorans ATCC 35296 AAC38572.2 O65987  
 Clocl_1056   [Clostridium] clariflavum DSM 19732 AEV67733.1    
 Clocl_1566   [Clostridium] clariflavum DSM 19732 AEV68202.1    
 Clocl_1567   [Clostridium] clariflavum DSM 19732 AEV68203.1    
 Clocl_1806   [Clostridium] clariflavum DSM 19732 AEV68415.1    
 Clocl_1864   [Clostridium] clariflavum DSM 19732 AEV68472.1    
 Clocl_1975   [Clostridium] clariflavum DSM 19732 AEV68577.1    
 endoglucanase (Clocl_2225) 3.2.1.4 [Clostridium] clariflavum DSM 19732 AEV68817.1    
 Clocl_3001   [Clostridium] clariflavum DSM 19732 AEV69539.1    
 Clocl_3029   [Clostridium] clariflavum DSM 19732 AEV69564.1    
 Clocl_3038   [Clostridium] clariflavum DSM 19732 AEV69573.1    
 Clocl_3253   [Clostridium] clariflavum DSM 19732 AEV69768.1    
 Clocl_3917   [Clostridium] clariflavum DSM 19732 AEV70357.1    
 Clocl_3255   [Clostridium] clariflavum DSM 19732 AEV69770.1    
 Cel9E (Cel9E)   Clostridium josui BAI52932.1 D1MXA1  
 Cel9D (Cel9D)   Clostridium josui BAI52930.1 D1MX99  
 Cel9A (Cel9A)   Clostridium josui BAI52926.1 D1MX95  
 Cel9B (Cel9B)   Clostridium josui BAI52927.1 D1MX96  
 Cel9C (Cel9C)   Clostridium josui BAI52929.1 D1MX98  
 Clole_0038   Clostridium lentocellum DSM 5427 ADZ81800.1    
 Clole_0485   Clostridium lentocellum DSM 5427 ADZ82224.1    
 Clole_0683   Clostridium lentocellum DSM 5427 ADZ82416.1    
 Clole_0709   Clostridium lentocellum DSM 5427 ADZ82442.1    
 Clole_3667   Clostridium lentocellum DSM 5427 ADZ85350.1    
 Clole_0675 (fragment)   Clostridium lentocellum DSM 5427 ADZ82408.1    
 Closa_2958   [Clostridium] saccharolyticum WM1 ADL05492.1 D9R760  
 Cspa_c18610   Clostridium saccharoperbutylacetonicum N1-4(HMT) AGF55631.1    
 Cspa_c21400 (CeD)   Clostridium saccharoperbutylacetonicum N1-4(HMT) AGF55906.1    
 Cspa_c18590 (CelK1)   Clostridium saccharoperbutylacetonicum N1-4(HMT) AGF55629.1    
 Cspa_c53100 (Celk2) (fragment)   Clostridium saccharoperbutylacetonicum N1-4(HMT) AGF59055.1    
 CLSAP_19670 (CelD)   Clostridium saccharoperbutylacetonicum N1-504 AQR94654.1    
 CLSAP_50570 (CelK) (fragment)   Clostridium saccharoperbutylacetonicum N1-504 AQR97724.1    
 Clo1100_1827   Clostridium sp. BNL1100 AEY66037.1    
 Clo1100_1824   Clostridium sp. BNL1100 AEY66034.1    
 Clo1100_1825   Clostridium sp. BNL1100 AEY66035.1    
 Clo1100_1850   Clostridium sp. BNL1100 AEY66060.1    
 Clo1100_1958   Clostridium sp. BNL1100 AEY66152.1    
 Clo1100_1821   Clostridium sp. BNL1100 AEY66031.1    
 Clo1100_1822   Clostridium sp. BNL1100 AEY66032.1    
 Clo1100_0253   Clostridium sp. BNL1100 AEY64541.1    
 Clo1100_0580   Clostridium sp. BNL1100 AEY64857.1    
 Clo1100_0933   Clostridium sp. BNL1100 AEY65189.1    
 Clo1100_1851   Clostridium sp. BNL1100 AEY66061.1    
 Clo1100_1852   Clostridium sp. BNL1100 AEY66062.1    
 Clo1100_1990   Clostridium sp. BNL1100 AEY66182.1    
 Clo1100_2671   Clostridium sp. BNL1100 AEY66832.1    
 cellulase (CbhA) (fragment)   Clostridium sp. JC3 BAC65236.1 Q845W7  
 SAMN02745906_2621   [Clostridium] sphenoides JCM 1415 ATCC 19403 SET87522.1    
 endo-β-1,4-glucanase Z / avicelase I (CelZ) 3.2.1.4 Clostridium stercorarium CAA39010.1 P23659  
 Cst_c18960 (CelZ)   [Clostridium] stercorarium subsp. stercorarium DSM 8532 AGI39870.1
AGC68873.1
   
 CSTERTH_09150   [Clostridium] stercorarium subsp. thermolacticum DSM 2910 ANW99179.1    
 CPS_4776   Colwellia psychrerythraea 34H AAZ26299.1 Q47UV2  
 CMT41_16995   Colwellia sp. MT41 ALO36242.1    
 CCU_17750   Coprococcus sp. ART55/1 CBK83282.1 D5HEE9  
 CCU_24920   Coprococcus sp. ART55/1 CBK83841.1 D5HG08  
 A9D14_03440   Croceicoccus marinus E4A9 ARU17213.1    
 A9D14_01615   Croceicoccus marinus E4A9 ARU17159.1    
 A9D14_10990   Croceicoccus marinus E4A9 ARU16610.1    
 Cyagr_2021   Cyanobium gracile PCC 6307 AFY29144.1    
 CYFUS_008670   Cystobacter fuscus DSM 52655 ATB43190.1    
 CYFUS_007541   Cystobacter fuscus DSM 52655 ATB42065.1    
 CHU_0778 (Cel9E)   Cytophaga hutchinsonii ATCC 33406 ABG58065.1 Q11X01  
 CHU_0961   Cytophaga hutchinsonii ATCC 33406 ABG58238.1 Q11WH8  
 periplasmic endoglucanase 9C (CHU_1280) (Cel9C) 3.2.1.4 Cytophaga hutchinsonii ATCC 33406 ABG58552.1 Q11VL4  
 CHU_1335 (Cel9B)   Cytophaga hutchinsonii ATCC 33406 ABG58607.1 Q11VF9  
 endoglucanase 9A (CHU_1336) (Cel9A) 3.2.1.4 Cytophaga hutchinsonii ATCC 33406 ABG58608.1 Q11VF8  
 CHU_1655 (Cel9D)   Cytophaga hutchinsonii ATCC 33406 ABG58923.1 Q11UJ1  
 CHU_2235 (Cel9F)   Cytophaga hutchinsonii ATCC 33406 ABG59498.1 Q11SW6  
 SY84_11555   Deinococcus soli Cha et al. 2016 N5 AKH18527.1    
 FH5T_02390   Draconibacterium orientale FH5 AHW58823.1    
 FH5T_03385   Draconibacterium orientale FH5 AHW58962.1    
 FH5T_04820   Draconibacterium orientale FH5 AHW59138.1    
 FH5T_09915   Draconibacterium orientale FH5 AHW59812.1    
 FH5T_15785   Draconibacterium orientale FH5 AHW60607.1    
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