Glycoside Hydrolase Family 9

Activities in Familyendoglucanase (EC 3.2.1.4); endo-β-1,3(4)-glucanase / lichenase-laminarinase (EC 3.2.1.6); β-glucosidase (EC 3.2.1.21); lichenase / endo-β-1,3-1,4-glucanase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); cellobiohydrolase (EC 3.2.1.91); xyloglucan-specific endo-β-1,4-glucanase / endo-xyloglucanase (EC 3.2.1.151); exo-β-glucosaminidase (EC 3.2.1.165)
Mechanism Inverting
3D Structure Status( α / α ) 6
Catalytic Nucleophile/BaseAsp (experimental)
Catalytic Proton DonorGlu (experimental)
Noteformerly known as cellulase family E.
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Statistics GenBank accession (3721); Uniprot accession (679); PDB accession (46); 3D entries (19); cryst (0)
All (2785) Archaea (7) Bacteria (1612) Eukaryota (954) unclassified (212) Structure (19) Characterized (171)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | ... | 17 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D
 AKO66_11755   Bacillus pumilus NJ-V ANY97359.1    
 AMR71_11750   Bacillus pumilus NJ-V2 ALM45878.1    
 BEN31_08005   Bacillus pumilus PDSLzg-1 AOC56746.1    
 endo-β-1,4-glucanase 3.2.1.4 Bacillus pumilus S-27 AAF15367.1 Q9R9H6  
 BPUM_1558   Bacillus pumilus SAFR-032 ABV62240.1 A8FDC3  
 UP15_08375   Bacillus pumilus SH-B11 AMM88974.1    
 UP12_08090   Bacillus pumilus SH-B9 AMM97307.1    
 BTUAT1_15740 (BglC)   Bacillus pumilus TUAT1 BAT48708.1    
 VT48_08305   Bacillus pumilus W3 AKC66020.1    
 endo-1,4-β-glucanase (EgL)   Bacillus pumilus Z14 AGO86843.1    
 BRL64_08205   Bacillus safensis BRM1 ARD56162.1    
 RS87_08135   Bacillus safensis FO-36b AWI36696.1    
 BSL056_08085   Bacillus safensis KCTC 12796BP APJ10917.1    
 BSA145_16055   Bacillus safensis U14-5 APT47248.1    
 BSA171_11320   Bacillus safensis U17-1 APT54136.1    
 BSA41_06395   Bacillus safensis U41 APT49588.1    
 S101395_03072   Bacillus sonorensis SRCM101395 ASB89579.1    
 endo-β-1,4-glucanase IV 3.2.1.4 Bacillus sp. (in: Bacteria) KSM-522 BAA24918.1 P28622  
 CJO35_09275   Bacillus sp. 1s-1 ASV15338.1    
 endoglucanase A (Ega;Bα-EGA) 3.2.1.4 Bacillus sp. AC-1 ABM68635.1 A6Y862  
 BSZ43_09205   Bacillus sp. H15-1 APJ26958.1    
 QR42_07950   Bacillus sp. WP8 AIZ60200.1    
 BK049_00285   Bacillus xiamenensis VV3 AOZ87246.1    
 cellulase (CelA20) 3.2.1.4 bacterium enrichment culture clone CelA20 ACR23658.1 C5HG55  
 BRDCF_03830   Bacteroidales bacterium CF AGX07793.1
AGY53377.1
   
 BcellWH2_01410 (Celd_2)   Bacteroides cellulosilyticus WH2 ALJ58671.1    
 BcellWH2_02910   Bacteroides cellulosilyticus WH2 ALJ60149.1    
 BcellWH2_01356 (Celd_1)   Bacteroides cellulosilyticus WH2 ALJ58617.1    
 BUN20_13550   Bacteroides fragilis Q1F2 AUI47497.1    
 Bache_3234   Bacteroides helcogenes P 36-108 ADV45158.1    
 C3V43_11895   Bacteroides heparinolyticus F0111 AVM59063.1    
 C3V43_09685 (fragment)   Bacteroides heparinolyticus F0111 AVM57995.1    
 C3V43_09680   Bacteroides heparinolyticus F0111 AVM57994.1    
 endo-xyloglucanase (CelD;BoGH9A;BACOVA_02649;Bovatus_03068)   Bacteroides ovatus ATCC 8483 ALJ47676.1    
 BACOV975_00830   Bacteroides ovatus V975 SCV07074.1    
 C4H11_08425   Bacteroides zoogleoformans ATCC 33285 AVM52961.1    
 C4H11_08430 (fragment)   Bacteroides zoogleoformans ATCC 33285 AVM54034.1    
 GH9   Bacteroidetes bacterium AC2a AIJ19565.1    
 BZG35_07265   Brevundimonas sp. LM2 AQR61472.1    
 BZG35_00230   Brevundimonas sp. LM2 AQR60256.1    
 CEP68_00850   Brevundimonas vesicularis FDAARGOS_289 ASE38172.1    
 BKD03_00100   Brucella sp. 09RB8471 APX67955.1    
 BKD02_12255   Brucella sp. 09RB8910 APY15166.1    
 BR10RB9215_C20833   Brucella sp. 10RB9215 SBW16163.1    
 BR141012304_20837   Brucella sp. 141012304 SCD25303.1    
 BR141012304_20837   Brucella sp. 141012304 SCD25303.1    
 CIY_10500 (fragment)   Butyrivibrio fibrisolvens 16/4 CBK73904.1 D4J1B5  
 CIY_16470 (fragment)   Butyrivibrio fibrisolvens 16/4 CBK74409.1 D4J2S0  
 endo-1,4-glucanase C (Ced1) 3.2.1.4 Butyrivibrio fibrisolvens H17C CAA39264.1 P23658  
 bhn_I2617 (fragment)   Butyrivibrio hungatei MB2003 AOZ97649.1    
 bhn_I0834   Butyrivibrio hungatei MB2003 AOZ95868.1    
 bpr_I1593   Butyrivibrio proteoclasticus B316 ADL34330.1 E0RV60  
 bpr_I1500   Butyrivibrio proteoclasticus B316 ADL34237.1 E0RWD9  
 bpr_I2796 (fragment)   Butyrivibrio proteoclasticus B316 ADL35526.1 E0RZW0  
 cellulase (CelA;CbCelA;Athe_1867;Cbes1867;Cbes_1867;CbCBH48A) (Cel9A+Cbh48A) 3.2.1.176
3.2.1.4
Caldicellulosiruptor bescii DSM 6725 ACM60955.1
CAB06786.1
B9MKU7
P96311
4DOD[A]
4DOE[A]
 bifunctional cellulase / β-mannanase (CelC-ManB;CbCel9B/Man5A;Athe_1865;Cbes_1865;Orf1952) (Cel9B-Man5A) 3.2.1.4
3.2.1.78
Caldicellulosiruptor bescii DSM 6725 ACM60953.1 B9MKU5  
 endo-β-1,3-1,4-glucanase (Cel19;Calkr_2522) 3.2.1.73 Caldicellulosiruptor kristjanssonii I77R1B ADQ41951.1 E4S8G9  
 Calkro_0850   Caldicellulosiruptor kronotskyensis 2002 ADQ45727.1 E4SDB8  
 Calkro_0855 (fragment)   Caldicellulosiruptor kronotskyensis 2002 ADQ45731.1 E4SDC2  
 Calla_2385   Caldicellulosiruptor lactoaceticus 6A AEM74912.1    
 Calla_0018   Caldicellulosiruptor lactoaceticus 6A AEM72714.1    
 COB47_1669   Caldicellulosiruptor obsidiansis OB47 ADL42950.1 D9TFI0  
 COB47_1673   Caldicellulosiruptor obsidiansis OB47 ADL42953.1 D9TFI3  
 cellulase (CelA) 3.2.1.4 Caldicellulosiruptor saccharolyticus AAA91086.1 P22534  
 Csac_1076 (CelA)   Caldicellulosiruptor saccharolyticus DSM 8903 ABP66690.1 A4XIF5  
 Csac_1079 (CelC)   Caldicellulosiruptor saccharolyticus DSM 8903 ABP66693.1 A4XIF8  
 multidomain endoglucanase (CelE)   Caldicellulosiruptor sp. Tok7B.1 AAK06394.1 Q9AQG4  
 Cabys_1959   Caldithrix abyssi DSM 13497 LF13 APF18708.1    
 NIES267_18330   Calothrix parasitica NIES-267 BAY82354.1    
 Acid345_0426   Candidatus Koribacter versatilis Ellin345 ABF39431.1 Q1IUL9  
 Acid345_2677   Candidatus Koribacter versatilis Ellin345 ABF41678.1 Q1IN72  
 Acid_0198   Candidatus Solibacter usitatus Ellin6076 ABJ81212.1 Q02CK4  
 Acid_2606   Candidatus Solibacter usitatus Ellin6076 ABJ83595.1 Q024I1  
 Caci_0460   Catenulispora acidiphila DSM 44928 ACU69412.1 C7PX61  
 Caci_5258   Catenulispora acidiphila DSM 44928 ACU74117.1 C7Q7V0  
 AQ619_10845   Caulobacter henricii CB4 ALL13792.1    
 CSW64_16290   Caulobacter mirabilis FWC 38 ATQ43844.1    
 Cseg_3644   Caulobacter segnis ATCC 21756 ADG12067.1 D5VNJ4  
 B7G68_18650   Caulobacter segnis TK0059 AVQ03678.1    
 CA607_12115   Caulobacter vibrioides CB1 ATC29091.1    
 CA606_08845   Caulobacter vibrioides CB13b1a ATC32447.1    
 CC2227 / CC_2227   Caulobacter vibrioides CB15 AAK24198.1
NP_421030.1
Q9A667  
 CA608_11900   Caulobacter vibrioides CB2 ATC25179.1    
 CCNA_02310   Caulobacter vibrioides NA1000 ACL95775.1 B8GYK2  
 CO704_03665   Cedecea neteri FDAARGOS_392 ATF91249.1    
 LH23_01035   Cedecea neteri M006 AIR59285.1    
 endo-β-1,4-glucanase B (CenB;Celf_0019) (Cel9A) 3.2.1.4 Cellulomonas fimi ATCC 484 AAA23086.1
AEE44171.1
P26225  
 endo-β-1,4-glucanase C (CenC) (Cel9B) 3.2.1.4 Cellulomonas fimi ATCC 484 CAA40993.1
AAA23088.1
AEE45671.1
P14090  
 Celf_0045   Cellulomonas fimi ATCC 484 AEE44196.1    
 Celf_1481 (probable fragment)   Cellulomonas fimi ATCC 484 AEE45616.1    
 Celf_1705   Cellulomonas fimi ATCC 484 AEE45837.1    
 endoglucanase (CBP105) (processive) 3.2.1.4 Cellulomonas flavigena CDBB-531 AAW62376.2 Q3ZMA8  
 Cfla_1515   Cellulomonas flavigena DSM 20109 ADG74413.1 D5UD76  
 Cfla_3563   Cellulomonas flavigena DSM 20109 ADG76435.1
ARU79882.1
D5UDH9  
 Cfla_0016   Cellulomonas flavigena DSM 20109 ADG72936.1 D5UFH7  
 Cfla_0139   Cellulomonas flavigena DSM 20109 ADG73058.1 D5UG75  
 Cfla_3031   Cellulomonas flavigena DSM 20109 ADG75914.1 D5UL32  
 Celgi_0019   Cellulomonas gilvus ATCC 13127 AEI10553.1    
 Celgi_2074   Cellulomonas gilvus ATCC 13127 AEI12575.1    
 CBP52_02935   Cellulomonas sp. PSBB021 ASR54271.1    
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