Glycoside Hydrolase Family 5

Activities in Familyendo-β-1,4-glucanase / cellulase (EC 3.2.1.4); endo-β-1,4-xylanase (EC 3.2.1.8); β-glucosidase (EC 3.2.1.21); β-mannosidase (EC 3.2.1.25); β-glucosylceramidase (EC 3.2.1.45); glucan β-1,3-glucosidase (EC 3.2.1.58); licheninase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); glucan endo-1,6-β-glucosidase (EC 3.2.1.75); mannan endo-β-1,4-mannosidase (EC 3.2.1.78); cellulose β-1,4-cellobiosidase (EC 3.2.1.91); steryl β-glucosidase (EC 3.2.1.104); endoglycoceramidase (EC 3.2.1.123); chitosanase (EC 3.2.1.132); β-primeverosidase (EC 3.2.1.149); xyloglucan-specific endo-β-1,4-glucanase (EC 3.2.1.151); endo-β-1,6-galactanase (EC 3.2.1.164); hesperidin 6-O-α-L-rhamnosyl-β-glucosidase (EC 3.2.1.168); β-1,3-mannanase (EC 3.2.1.-); arabinoxylan-specific endo-β-1,4-xylanase (EC 3.2.1.-); mannan transglycosylase (EC 2.4.1.-)
Mechanism Retaining
ClanGH-A
3D Structure Status( β / α ) 8
Catalytic Nucleophile/BaseGlu (experimental)
Catalytic Proton DonorGlu (experimental)
NoteOnce known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)MEGAZYME; NZYTech; PROZOMIX;
Statistics GenBank accession (11393); Uniprot accession (1922); PDB accession (208); 3D entries (73); cryst (0)
All (10017) Archaea (67) Bacteria (7846) Eukaryota (1982) Viruses (5) unclassified (117) Structure (73) Characterized (547)
| 1 | ... | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 10 | 11 | ... | 20 |
Eukaryota
Protein Name EC#OrganismGenBank UniprotPDB/3D Subf
 MG05844.4   Magnaporthe oryzae 70-15 XP_369620.1     7
 MG06512.4   Magnaporthe oryzae 70-15 XP_369997.1 G4N6W3   9
 MG00319.4   Magnaporthe oryzae 70-15 XP_368925.1 G4NCZ5   5
 MG00530.4   Magnaporthe oryzae 70-15 XP_368714.1 G4NBI5   24
 MG00882.4   Magnaporthe oryzae 70-15 XP_368362.1 G4NDR4   12
 MG03374.4   Magnaporthe oryzae 70-15 XP_360831.1 G4N8V8   16
 MG05381.4   Magnaporthe oryzae 70-15 XP_360006.1 G4MKY3   16
 MG09342.4 (fragment)   Magnaporthe oryzae 70-15 XP_364584.1 G4NI55   7
 MG10083.4   Magnaporthe oryzae 70-15 XP_365863.1 G4N908   5
 MG09433.4   Magnaporthe oryzae 70-15 XP_364493.1     22
 MGG_06402   Magnaporthe oryzae 70-15 XP_003717168.1
XP_369887.1
    11
 MGG_10423   Magnaporthe oryzae 70-15 XP_003719440.1
XP_366204.1
    16
 MGL_0748   Malassezia globosa CBS 7966 EDP44941.1     50
 MSYG_3070   Malassezia sympodialis ATCC 42132 SHO78723.1     50
 MSYG_2409 (fragment)   Malassezia sympodialis ATCC 42132 SHO78067.1     12
 MSYG_2412   Malassezia sympodialis ATCC 42132 SHO78070.1     50
 MSYG_4240   Malassezia sympodialis ATCC 42132 SHO79887.1     50
 MSYG_4116   Malassezia sympodialis ATCC 42132 SHO79766.1     9
 MSYG_3905   Malassezia sympodialis ATCC 42132 SHO79555.1     50
 MSYG_3512 (fragment)   Malassezia sympodialis ATCC 42132 SHO79163.1     9
 endo-β-mannanase (Man2;MdMAN2)   Malus domestica ACN78664.1 C7A7X8   7
 endo-β-mannanase (Man3;MdMAN3)   Malus domestica ACN78665.1 C7A7X9   7
 endo-β-mannanase (Man1;MdMAN1)   Malus domestica ACN78663.1 C7A7X7   7
 glycoside hydrolase family 5 endoglucanase, partial (fragment)   Marasmius oreades FB-MO-71 CDL74348.1     5
 unknown   Medicago truncatula AFK33638.1     14
 unknown   Medicago truncatula AFK42423.1     14
 MtrDRAFT_AC149642g20v1   Medicago truncatula ABD32354.1 Q2HTU0   11
 MtrDRAFT_AC145165g35v2   Medicago truncatula ABP02833.1
ABE91798.1
ABE91799.1
    14
 MtrDRAFT_AC137669g17v2   Medicago truncatula ABO79647.1     7
 MtrDRAFT_AC141435g4v2 + MtrDRAFT_AC141435g5v2 (fragments)   Medicago truncatula ABO80563.1
ABO80564.1
    11
 MtrDRAFT_AC147959g9v1   Medicago truncatula ABP03047.1
AFK37837.1
    14
 MtrDRAFT_AC147959g24v1   Medicago truncatula ABP03049.1     14
 MtrDRAFT_AC147959g10v1   Medicago truncatula ABP03050.1     14
 BN887_02050   Melanopsichium pennsylvanicum 4 CDI51178.1     9
 BN887_01986   Melanopsichium pennsylvanicum 4 CDI51074.1     50
 BN887_02040   Melanopsichium pennsylvanicum 4 CDI51196.1     12
 BN887_00859   Melanopsichium pennsylvanicum 4 CDI51800.1     9
 BN887_05784 (fragment)   Melanopsichium pennsylvanicum 4 CDI52106.1     50
 BN887_04738   Melanopsichium pennsylvanicum 4 CDI54550.1     12
 BN887_04978   Melanopsichium pennsylvanicum 4 CDI55234.1     9
 BN887_04322   Melanopsichium pennsylvanicum 4 CDI55906.1     16
 BN887_01891   Melanopsichium pennsylvanicum 4 CDI56177.1     9
 BN887_03423   Melanopsichium pennsylvanicum 4 CDI56700.1    
 Mard-eng-1 (fragment)   Meloidogyne ardenensis AER27760.1     2
 β-1,4-endoglucanase-1, partial (Eng-1) (fragment)   Meloidogyne arenaria AHX24636.1     2
 β-1,4-endoglucanase Eng-1ii (fragment)   Meloidogyne arenaria AAK21893.1 Q9BJY3   2
 β-1,4-endoglucanase Eng-1i (fragment)   Meloidogyne arenaria AAK21892.1 Q9BJY4   2
 β-1,4-endoglucanase (fragment)   Meloidogyne artiellia AER27761.1     2
 β-1,4-endoglucanase (fragment)   Meloidogyne chitwoodi AER27805.1     2
 β-1,4-endoglucanase (fragment)   Meloidogyne chitwoodi AER27807.1     2
 endo-β-1,4-glucanase   Meloidogyne chitwoodi Mc-eng-2 AER27806.1     2
 β-1,4-endoglucanase (fragment)   Meloidogyne hapla AAP33282.1 Q86BU9  
 β-1,4-endoglucanase-1, partial (Eng-1) (fragment)   Meloidogyne hapla AHX24637.1     2
 β-1,4-endoglucanase Eng-1i (fragment)   Meloidogyne hapla AAK21891.1
AAS48572.1
Q9BJY5   2
 β-1,4-endoglucanase-1, partial (Eng-1) (fragment)   Meloidogyne hispanica AHX24640.1     2
 β-1,4-endoglucanase (fragment)   Meloidogyne ichinohei AER27762.1     2
 β-1,4-endoglucanase (fragment)   Meloidogyne ichinohei AER27763.1     2
 endo-β-1,4-glucanase 1 (Eng-1) 3.2.1.4 Meloidogyne incognita AAD45868.1 Q9UA57   2
 cellulase (Eng4)   Meloidogyne incognita AAR37375.1 Q5QJ65   2
 cellulase (Eng3;Eng-1)   Meloidogyne incognita AAR37374.1
AHX24639.1
Q5QJ66   2
 β-1,4-endoglucanase Eng-1vi (fragment)   Meloidogyne incognita AAK21890.1 Q9BH59   2
 β-1,4-endoglucanase Eng-1v (fragment)   Meloidogyne incognita AAK21889.1 Q9BJY6   2
 β-1,4-endoglucanase Eng-1iv (fragment)   Meloidogyne incognita AAK21888.1 Q9BH59   2
 β-1,4-endoglucanase Eng-1iii (fragment)   Meloidogyne incognita AAK21887.1 Q9BH59   2
 β-1,4-endoglucanase Eng-1ii (fragment)   Meloidogyne incognita AAK21886.1 Q9BJY7   2
 β-1,4-endoglucanase Eng-1i (fragment)   Meloidogyne incognita AAK21885.1 Q9BJY8   2
 β-1,4-endoglucanase Eng-2b (fragment)   Meloidogyne incognita AAK21884.1 Q9BJY9  
 β-1,4-endoglucanase (Eng-2) 3.2.1.4 Meloidogyne incognita AAK21883.2 Q9BJZ0   2
 endo-β-1,4-glucanase (Eng-1A;Mi-eng-1) 3.2.1.4 Meloidogyne incognita AAK21882.1 Q9BJZ1   2
 β-1,4-endoglucanase (Eng-2) 3.2.1.4 Meloidogyne incognita AAK21881.1 Q9BJZ2   2
 β-1,4-endoglucanase-1, partial (Eng-1) (fragment)   Meloidogyne javanica AHX24638.1     2
 β-1,4-endoglucanase Eng-1ii (fragment)   Meloidogyne javanica AAK21895.1 Q9BJY1   2
 β-1,4-endoglucanase Eng-1i (fragment)   Meloidogyne javanica AAK21894.1 Q9BJY2   2
 cellulase (Eng-3)   Meloidogyne javanica HYN CAJ77137.1
ACA66271.1
A2VA30   2
 cellulase 5 (Cgh5A)   Mesosa myops AMA76414.1     2
 cellulase 5 (Cgh5D)   Mesosa myops AMA76418.1     2
 cellulase 5 (Cgh5F)   Mesosa myops AMA76419.1     2
 cellulase 5, partial (Cgh5G) (fragment)   Mesosa myops AMA76420.1    
 cellulase 5, partial (Cgh5E)   Mesosa myops AMA76417.1     2
 cellulase 5 (Cgh5B)   Mesosa myops AMA76415.1     2
 cellulase 5 (Cgh5C)   Mesosa myops AMA76416.1     2
 MAC_09197   Metarhizium acridum CQMa 102 EFY84753.1     15
 endo-β-mannanase   Metopus es CCAP 1653/2 ABR27262.1 A6MI38   41
 exo-β-1,3-glucanase   Meyerozyma caribbica AEP83833.1     9
 exo-β-1,3-glucanase (Exg1)   Meyerozyma guilliermondii K123-1 / M8 ACN18104.1
ADV90770.1
A5DN91   9
 exo-β-1,3-glucanase (fragment)   Meyerozyma guilliermondii M8 ABY47601.1 A9YX59   9
 ORF (fragment)   Microdochium bolleyi 0629-022 2 WA/MF06-285 CDL74339.1     5
 MICPUN_58085   Micromonas commoda sp. RCC299 ACO62742.1 C1E4L9   7
 MICPUN_83090 (fragment)   Micromonas commoda sp. RCC299 ACO64669.1 C1E9E6   7
 MICPUN_102881   Micromonas commoda sp. RCC299 ACO69881.1 C1FHU7  
 GNLVRS01_PISO0A03784g (Piso0_000184)   Millerozyma farinosa CBS 7064 CCE72597.1     9
 GNLVRS01_PISO0B03917g (Piso0_000184)   Millerozyma farinosa CBS 7064 CCE73161.1     9
 GNLVRS01_PISO0N05909g (Piso0_005010)   Millerozyma farinosa CBS 7064 CCE86515.1     49
 GNLVRS01_PISO0M05754g (Piso0_005010)   Millerozyma farinosa CBS 7064 CCE85414.1     49
 GNLVRS01_PISO0L01419g (Piso0_003735)   Millerozyma farinosa CBS 7064 CCE84194.1     12
 GNLVRS01_PISO0K01418g (Piso0_003735)   Millerozyma farinosa CBS 7064 CCE83163.1     12
 GNLVRS01_PISO0D12411g (Piso0_001166)   Millerozyma farinosa CBS 7064 CCE79126.1     9
 GNLVRS01_PISO0D12587g (Piso0_001174)   Millerozyma farinosa CBS 7064 CCE79134.1     9
 GNLVRS01_PISO0C12344g (Piso0_001166)   Millerozyma farinosa CBS 7064 CCE78540.1     9
 GNLVRS01_PISO0C12520g (Piso0_001174)   Millerozyma farinosa CBS 7064 CCE78548.1     9
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