Glycoside Hydrolase Family 5

Activities in Familyendo-β-1,4-glucanase / cellulase (EC 3.2.1.4); endo-β-1,4-xylanase (EC 3.2.1.8); β-glucosidase (EC 3.2.1.21); β-mannosidase (EC 3.2.1.25); β-glucosylceramidase (EC 3.2.1.45); glucan β-1,3-glucosidase (EC 3.2.1.58); licheninase (EC 3.2.1.73); exo-β-1,4-glucanase / cellodextrinase (EC 3.2.1.74); glucan endo-1,6-β-glucosidase (EC 3.2.1.75); mannan endo-β-1,4-mannosidase (EC 3.2.1.78); cellulose β-1,4-cellobiosidase (EC 3.2.1.91); steryl β-glucosidase (EC 3.2.1.104); endoglycoceramidase (EC 3.2.1.123); chitosanase (EC 3.2.1.132); β-primeverosidase (EC 3.2.1.149); xyloglucan-specific endo-β-1,4-glucanase (EC 3.2.1.151); endo-β-1,6-galactanase (EC 3.2.1.164); hesperidin 6-O-α-L-rhamnosyl-β-glucosidase (EC 3.2.1.168); β-1,3-mannanase (EC 3.2.1.-); arabinoxylan-specific endo-β-1,4-xylanase (EC 3.2.1.-); mannan transglycosylase (EC 2.4.1.-)
Mechanism Retaining
ClanGH-A
3D Structure Status( β / α ) 8
Catalytic Nucleophile/BaseGlu (experimental)
Catalytic Proton DonorGlu (experimental)
NoteOnce known as cellulase family A; many members have been assigned to subfamilies as described by Aspeborg et al. (2012) BMC Evol Biol. 12(1):186 (PMID: 22992189).
External resourcesCAZypedia; HOMSTRAD; PROSITE;
Commercial Enzyme Provider(s)MEGAZYME; NZYTech; PROZOMIX;
Statistics GenBank accession (12681); Uniprot accession (1922); PDB accession (213); 3D entries (75); cryst (0)
All (11302) Archaea (67) Bacteria (9025) Eukaryota (2084) Viruses (6) unclassified (120) Structure (75) Characterized (547)
| 1 | ... | 12 | 13 | 14 | 15 | 16 | 17 | 18 | 19 | 20 | 21 |
Eukaryota
Protein Name EC#OrganismGenBank UniprotPDB/3D Subf
 NT0285B14   uncultured symbiotic protist of Hodotermopsis sjoestedti BAF57330.1 A4UWS0   2
 NT0285B44   uncultured symbiotic protist of Hodotermopsis sjoestedti BAF57331.1 A4UWS1   1
 NT0287A37;NT0290A76;NT0290B18;NT0290B53   uncultured symbiotic protist of Hodotermopsis sjoestedti BAF57332.1
BAF57336.1
BAF57338.1
BAF57339.1
A4UWS2   2
 NT0287A89   uncultured symbiotic protist of Hodotermopsis sjoestedti BAF57333.1 A4UWS3   4
 NT0288A58   uncultured symbiotic protist of Hodotermopsis sjoestedti BAF57334.1 A4UWS4   2
 NT0288B91   uncultured symbiotic protist of Hodotermopsis sjoestedti BAF57335.1 A4UWS5   1
 NT0291B62;NT0290A90;NT0291A24   uncultured symbiotic protist of Hodotermopsis sjoestedti BAF57341.1
BAF57337.1
BAF57340.1
A4UWT1   2
 SM2760A61   uncultured symbiotic protist of Mastotermes darwiniensis BAF57413.1 A4UX03   2
 SM2763A16   uncultured symbiotic protist of Mastotermes darwiniensis BAF57414.1 A4UX04   1
 SM2765A45   uncultured symbiotic protist of Mastotermes darwiniensis BAF57415.1 A4UX05   1
 SM2632A46;SM2637A04   uncultured symbiotic protist of Neotermes koshunensis BAF57361.1
BAF57367.1
A4UWV1   2
 SM2632A58   uncultured symbiotic protist of Neotermes koshunensis BAF57362.1 A4UWV2   1
 SM2632B73;SM2637B70;SM2649A06;SM2649A59;SM2649B23   uncultured symbiotic protist of Neotermes koshunensis BAF57363.1
BAF57368.1
BAF57369.1
BAF57370.1
BAF57373.1
A4UWV3   2
 SM2635A05;SM2636A42;SM2649A74;SM2649A87   uncultured symbiotic protist of Neotermes koshunensis BAF57364.1
BAF57365.1
BAF57371.1
BAF57372.1
A4UWV4   1
 SM2636B47   uncultured symbiotic protist of Neotermes koshunensis BAF57366.1 A4UWV6   2
 SM2030A48   uncultured symbiotic protist of Reticulitermes speratus BAF57288.1 A4UWM8   1
 SM2040A21   uncultured symbiotic protist of Reticulitermes speratus BAF57289.1 A4UWM9   1
 SM2040A25   uncultured symbiotic protist of Reticulitermes speratus BAF57290.1 A4UWN0   2
 SM2041A08 (possible fragment)   uncultured symbiotic protist of Reticulitermes speratus BAF57291.1 A4UWN1   1
 SM2041A23   uncultured symbiotic protist of Reticulitermes speratus BAF57292.1 A4UWN2   2
 SM2041A77 (fragment)   uncultured symbiotic protist of Reticulitermes speratus BAF57293.1 A4UWN3   2
 UBRO_00370   Ustilago bromivora UB2112 SAM71258.1     9
 UBRO_00392   Ustilago bromivora UB2112 SAM71140.1     12
 UBRO_06823   Ustilago bromivora UB2112 SAM84188.1     9
 UBRO_08206   Ustilago bromivora UB2112 SAM85626.1     9
 UBRO_01770   Ustilago bromivora UB2112 SAM77391.1     50
 UBRO_08657   Ustilago bromivora UB2112 SAM86230.1    
 UBRO_00201   Ustilago bromivora UB2112 SAM72210.1     50
 UBRO_07788   Ustilago bromivora UB2112 SAM85236.1     16
 UBRO_01319   Ustilago bromivora UB2112 SAM75227.1     9
 UBRO_05824   Ustilago bromivora UB2112 SAM84544.1     12
 UM00128.1 (Umay-Bgl506)   Ustilago maydis 521 EAK81513.1     50
 endo-β-mannanase (Man1)   Vaccinium corymbosum ACN78666.1 C7A7Y0   7
 ORF   Vaucheria litorea AFA52581.1    
 β-1,6-endoglucanase (VegB)   Verticillium dahliae AGW80454.1     15
 ORF 1 (fragment)   Verticillium dahliae 76 AAX73408.1 Q56NK6   12
 β-1,6-glucanase (Bgn3) (fragment)   Verticillium dahliae 76 AAX73406.1 Q56NK8   15
 VIGAN_11175000 (Vigan.11g175000)   Vigna angularis var. angularis BAU02258.1     7
 VIGAN_07175900 (Vigan.07g175900)   Vigna angularis var. angularis BAT92896.1     14
 VIGAN_09003000 (Vigan.09g003000)   Vigna angularis var. angularis BAT96746.1     7
 VIGAN_09149100 (Vigan.09g149100)   Vigna angularis var. angularis BAT97907.1     11
 VIGAN_10160100 (Vigan.10g160100)   Vigna angularis var. angularis BAU00044.1     7
 VIGAN_06132000 (Vigan.06g132000)   Vigna angularis var. angularis BAT90136.1     7
 VIGAN_06158400 (Vigan.06g158400)   Vigna angularis var. angularis BAT90354.1     11
 VIGAN_01093100 (Vigan.01g093100)   Vigna angularis var. angularis BAT73448.1     11
 VIGAN_03283600 (Vigan.03g283600)   Vigna angularis var. angularis BAT82775.1     7
 VIGAN_UM166800 (Vigan.umg166800) (fragment)   Vigna angularis var. angularis BAU03734.1     11
 VIGAN_02034400 (Vigan.02g034400)   Vigna angularis var. angularis BAT77749.1     7
 VIGAN_01135500 (Vigan.01g135500) (fragment)   Vigna angularis var. angularis BAT73817.1     11
 VIGAN_04376600 (Vigan.04g376600)   Vigna angularis var. angularis BAT86142.1     7
 VIGAN_03283800 (Vigan.03g283800) (fragment)   Vigna angularis var. angularis BAT82777.1     7
 VIGAN_03157600 (Vigan.03g157600)   Vigna angularis var. angularis BAT81736.1     14
 VIGAN_04421400 (Vigan.04g421400)   Vigna angularis var. angularis BAT86547.1     14
 VIGAN_01093300 (Vigan.01g093300) (fragment)   Vigna angularis var. angularis BAT73449.1     11
 VIGAN_01084100 (Vigan.01g084100)   Vigna angularis var. angularis BAT73366.1     14
 VIGAN_03157700 (Vigan.03g157700) (fragment)   Vigna angularis var. angularis BAT81737.1     14
 VIGAN_04114500 (Vigan.04g114500)   Vigna angularis var. angularis BAT83903.1     7
 VIGAN_04380200 (Vigan.04g380200)   Vigna angularis var. angularis BAT86176.1     7
 ORF (fragment)   Vitis riparia AAS00040.1 Q6R4Q6  
 VITISV_020441 (short fragment)   Vitis vinifera CAN62216.1 A5C9G4   7
 VITISV_015565 (short fragment)   Vitis vinifera CAN71607.1 A5AS93   7
 VITISV_000586   Vitis vinifera CAN74231.1 A5B782   14
 VITISV_020726 (fragment)   Vitis vinifera CAN70632.1 A5BWB8   7
 VITISV_023481 (short fragment)   Vitis vinifera CAN71993.1 A5AS93   7
 VITISV_023482 (short fragment)   Vitis vinifera CAN71994.1 A5B3T1   7
 VITISV_023484   Vitis vinifera CAN71995.1 A5B3T2   7
 VITISV_016766 + VITISV_016765 (fragments)   Vitis vinifera CAN78585.1
CAN78586.1
    14
 VITISV_033630   Vitis vinifera CAN77937.1 A5AHI3   7
 EgL (fragment)   Volvariella bombycina AEQ58920.1     5
 endoglucanase I (EG1;EGI) 3.2.1.4 Volvariella volvacea V14 AAG59832.1
AEQ58914.1
Q9C3Z8   5
 endoglucanase I (EgI;NCel5A) (Cel5A) 3.2.1.4 Volvariella volvacea WX32 ACE06751.1     5
 exo-β-1,3-glucanase   Wickerhamomyces anomalus ARX76274.1     9
 exo-β-1,3-glucanase (Exg1;PaEXG1;WaEXG1)   Wickerhamomyces anomalus CAA05243.1 H6UWP0
O93939
  9
 exo-1,3-β-glucanase (Exg2;PaEXG2)   Wickerhamomyces anomalus AGC67022.1     9
 exo-β-1,3-glucanase (Exg1)   Wickerhamomyces anomalus AS F17.12 CDF77670.1     9
 exo-β-1,3-glucanase (Exg2)   Wickerhamomyces anomalus AS F17.12 CDF77671.1     9
 exo-β-1,3-glucanase (Exg2)   Wickerhamomyces anomalus AS1 CDH93665.1     9
 exo-β-1,3-glucanase (Exg2;WaEXG2)   Wickerhamomyces anomalus BCA15 AFK33204.1     9
 exo-β-1,3-glucanase (Exg1;WaEXG1) (fragment)   Wickerhamomyces anomalus BCA15 AFK33201.1     9
 exo-β-1,3-glucanase (Exg1;WaEXG1)   Wickerhamomyces anomalus BCU24 AFK33202.1     9
 exo-β-1,3-glucanase (Exg2;WaEXG2)   Wickerhamomyces anomalus BCU24 AFK33205.1     9
 exo-β-1,3-glucanase (Exg1;WaEXG1)   Wickerhamomyces anomalus BS91 AFK33200.1     9
 exo-β-1,3-glucanase (Exg2;WaEXG2)   Wickerhamomyces anomalus BS91 AFK33203.1     9
 exo-β-1,3-glucanase (Exg2;PaEXG2;WaEXG2) 3.2.1.58 Wickerhamomyces anomalus K / NCYC 432 CAA11018.1 O93983   9
 WAN_9541   Wickerhamomyces anomalus NRRL Y-366 AEZ66638.1     9
 WAN_6658   Wickerhamomyces anomalus NRRL Y-366 AEZ66635.1     9
 WAN_362   Wickerhamomyces anomalus NRRL Y-366 AEZ66636.1     9
 WAN_2708   Wickerhamomyces anomalus NRRL Y-366 AEZ66637.1     9
 WAN_5696   Wickerhamomyces anomalus NRRL Y-366 AEZ66639.1     49
 WAN_7786   Wickerhamomyces anomalus NRRL Y-366 AEZ66642.1     49
 WAN_7517   Wickerhamomyces anomalus NRRL Y-366 AEZ66640.1     12
 WAN_7540   Wickerhamomyces anomalus NRRL Y-366 AEZ66641.1     12
 exo-1,3-β-glucanase (Exg1;PaEXG1)   Wickerhamomyces anomalus WRL-076 AGC67021.1     9
 exo-β-1,3-glucanase (Exg2;PaEXG2;WaEXG2)   Wickerhamomyces anomalus YF07b ABK40520.1 A0MPR7   9
 exo-β-1,3-glucanase (Exg1;WsExg1) 3.2.1.58 Williopsis saturnus WC91-2 ACP74152.2
ACP74152.1
C3VMY4   9
 glycoside hydrolase   Xanthophyllomyces dendrorhous CDZ97258.1     12
 Glycoside hydrolase, superfamily   Xanthophyllomyces dendrorhous CDZ96181.1     50
 Glycoside hydrolase, superfamily   Xanthophyllomyces dendrorhous CDZ97301.1     12
 Glycoside hydrolase, superfamily   Xanthophyllomyces dendrorhous CED85016.1     12
 glycoside hydrolase   Xanthophyllomyces dendrorhous CDZ98740.1     50
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