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Glycoside Hydrolase Family 56

Activities in FamilyChondroitinase / chondroitin hydrolase (EC 3.2.1.-);
Hyaluronidase / hyaluronoglucosaminidase (EC 3.2.1.35);
Mechanism Retaining e
3D Structure Status( β / α ) 7 barrel
Catalytic Nucleophile/Basecarbonyl oxygen of C-2 acetamido group of substrate
Catalytic Proton DonorGlu
External resourcesCAZypedia; PRINTS;
Statistics GenBank accession (532); Uniprot accession (63); PDB accession (6); 3D entries (3); cryst (0)
Summary
Download GH56 (372) Taxonomic display Structure (3) Characterized (29)
| 1 | 2 | 3 | 4 |
Eukaryota
Protein Name EC#OrganismGenBank UniprotPDB/3D
 hyaluronidase   Ampulex compressa ARK19841.1    
 venom hyaluronidase, partial (fragment)   Androctonus bicolor AIX87602.1    
 hyaluronidase   Androctonus crassicauda QQS80959.1    
 hyaluronidase (Hase-1)   Androctonus crassicauda UCR74866.1    
 hyaluronidase (HyaL)   Anoplius samariensis BAF93867.1    
 hyaluronidase 2   Apis cerana cerana AAK51798.2    
 hyaluronidase (HYA;Api m 2;Apim2;LOC406146) 3.2.1.35 Apis mellifera AAA27730.1
AAB45920.1
AAR78242.1
ABH87698.1
NP_001011619.1		 Q08169 1FCQ[A]
1FCU[A]
1FCV[A]
2J88[A]
 hyaluronidase   Apis mellifera carnica AFI40555.1    
 ORF   Ascaris suum ADY46439.1    
 Hyal1 (fragment)   Balaenoptera acutorostrata AEB78445.1    
 Hyal1 (fragment)   Balaenoptera omurai ADZ57824.1    
 BEMITA_ LOCUS 10475   Bemisia tabaci CAH0391897.1    
 LolHyaz (fragment)   Bichromomyia olmeca ANW11472.1    
 hyaluronidase   Bitis arietans ABI33944.1
ABI33945.1
ABI33946.1
ABI33947.1
ABI33948.1		    
 PH-20 hyaluronidase   Blarina brevicauda QKV49747.1    
 hyaluronidase, partial   Boiga irregularis AWX67683.1    
 Hyal2   Bos taurus AAL78385.1
AAI02043.1
AAX46601.1
AAX46657.1
ABG66994.1
NP_776772.1		    
 hyaluronidase (Spam1;PH-20;p80;bovHyal) 3.2.1.35 Bos taurus AAI10184.1
AAP55713.1
AAV41022.1		 Q2YDK3
Q5U7L5
Q7YS45		  
 HYAL1 (fragment)   Bos taurus AAX08792.1    
 HYAL1   Bos taurus AAX09052.1
AAI02474.1		    
 hyaluronidase   Bothrops atrox AUF71538.1    
 Hyaluronidase BmooHyal-2, partial   Bothrops moojeni ATU85543.1    
 Hyaluronidase BmooHyal-1, partial   Bothrops moojeni ATU85542.1    
 hyaluronidase (Hyase;BpHyase) (fragment)   Bothrops neuwiedi pauloensis ACN58137.1    
 hyaluronidase (Hyal;BvHyal) (partial) 3.2.1.35 Brachypelma vagans AFS33217.1 J9XYC6  
 MELIAE_ LOCUS 2367 (fragment)   Brassicogethes aeneus CAH0549075.1    
 BINO364_ LOCUS 2122   Brenthis ino CAH0715151.1    
 hyaluronidase PH-20, partial (fragment)   Bubalus bubalis bubalis ANC49763.1    
 hyaluronidase PH-20, partial (fragment)   Bubalus bubalis bubalis ANC49764.1    
 L3Y34_017400   Caenorhabditis briggsae isolate QX1410_ONT ULU04609.1    
 hyaluronidase / chondroitinase (Hyα-1;T22C8.2) 3.2.1.-
3.2.1.35		 Caenorhabditis elegans Bristol N2 CAA88874.1
BAG48551.1
NP_495830.1		 G5ECE8
Q22675		  
 7969   Cafeteria roenbergensis BVI (Cafroe1)      
 7720   Cafeteria roenbergensis BVI (Cafroe1)      
 PH-20   Callimico goeldii ACV60225.1    
 PH-20 (fragment)   Callimico goeldii ABD94489.1    
 hyaluronidase   Calliophis bivirgatus QSI83992.1    
 HYAL3-like protein (fragment)   Callithrix jacchus ABQ22669.1    
 ORF   Callorhinchus milii AFO99844.1    
 ORF   Callorhinchus milii AFO97416.1    
 ORF   Callorhinchus milii AFP02624.1    
 Hyal1 (fragment)   Camelus bactrianus AEB78448.1    
 sperm membrane protein (PH-20) (fragment)   Capra hircus ABI31753.1    
 hyaluronidase PH-20   Cavia porcellus CAA39768.1
AAB45922.1		    
 hyaluronidase   Cerastes cerastes ABI33938.1
ABI33939.1
ABI33940.1		    
 ORF (fragment)   Ceratonova shasta UKS51845.1    
 Hyal1 (fragment)   Ceratotherium simum AEB78449.1    
 PH-20   Cercopithecus mitis ACV60221.1    
 PH-20 (fragment)   Cercopithecus mitis ABD94493.1    
 CEUTPL_ LOCUS 8307 (fragment)   Ceutorhynchus assimilis CAG9767750.1    
 CEUTPL_ LOCUS 8310 (fragment)   Ceutorhynchus assimilis CAG9767753.1    
 CEUTPL_ LOCUS 8308 (fragment)   Ceutorhynchus assimilis CAG9767751.1    
 CEUTPL_ LOCUS 8309 (fragment)   Ceutorhynchus assimilis CAH1129628.1    
 venom protein (A4YE08CM1) (fragment)   Chelonus inanitus CBN72517.1    
 venom protein (A3YM10CM1) (fragment)   Chelonus inanitus CBN72518.1    
 venom protein (A2YM17CM2) (fragment)   Chelonus inanitus CBN72519.1    
 venom protein (A1YB14CM3) (fragment)   Chelonus inanitus CBN72520.1    
 CHILSU_ LOCUS 6564   Chilo suppressalis CAH0403295.1    
 CHIRRI_ LOCUS 55   Chironomus riparius CAG9797054.1    
 CHIRRI_ LOCUS 4158   Chironomus riparius CAG9801227.1    
 posterior head protein 20, partial (Ph-20)   Chlorocebus aethiops AEX20366.1    
 CINC_ LOCUS 8013   Chrysodeixis includens CAH0598162.1    
 395682   Ciona intestinalis (Cioin2)      
 378054   Ciona intestinalis (Cioin2)      
 posterior head protein 20, partial (Ph-20)   Colobus guereza AEX20365.1    
 Conohyal-Ad1 (fragment)   Conus adamsonii AFH78529.1    
 hyaluronidase (Conohyal-Cn1) 3.2.1.35 Conus consors IV AFH78528.1 I0CME7  
 venom-related protein conohyaluronidase, partial   Conus ebraeus UMA82668.1    
 conohyaluronidase   Conus ermineus AXL95480.1    
 conohyaluronidase, partial (fragment)   Conus ermineus AXL95522.1    
 conohyaluronidase, partial (fragment)   Conus ermineus AXL95351.1    
 conohyaluronidase, partial (fragment)   Conus magus QFQ61159.1    
 conohyaluronidase   Conus magus QFQ61160.1    
 conohyaluronidase   Conus magus QFQ61157.1    
 conohyaluronidase   Conus magus QFQ61158.1    
 ORF (fragment)   Coptotermes formosanus AGM32945.1    
 CSTEINMANNI_LOCUS2772392 (fragment)   Coregonus sp. balchen CAB1336289.1    
 CSTEINMANNI_LOCUS2772377   Coregonus sp. balchen CAB1336274.1    
 CSTEINMANNI_LOCUS3956487   Coregonus sp. balchen CAB1345787.1    
 CSTEINMANNI_LOCUS2290463   Coregonus sp. balchen CAB1331596.1    
 CSTEINMANNI_LOCUS352570   Coregonus sp. balchen CAB1314663.1    
 CSTEINMANNI_LOCUS4885995 (fragment)   Coregonus sp. balchen CAB1353427.1    
 CSTEINMANNI_LOCUS1271682   Coregonus sp. balchen CAB1323488.1    
 CSTEINMANNI_LOCUS4886719   Coregonus sp. balchen CAB1354151.1    
 CSTEINMANNI_LOCUS2290462   Coregonus sp. balchen CAB1331595.1    
 CSTEINMANNI_LOCUS1271681   Coregonus sp. balchen CAB1323487.1    
 CSTEINMANNI_LOCUS2048482   Coregonus sp. balchen CAB1329445.1    
 CSTEINMANNI_LOCUS2048494   Coregonus sp. balchen CAB1329457.1    
 CSTEINMANNI_LOCUS353316   Coregonus sp. balchen CAB1315409.1    
 CSTEINMANNI_LOCUS1271683   Coregonus sp. balchen CAB1323489.1    
 CSTEINMANNI_LOCUS4885994   Coregonus sp. balchen CAB1353426.1    
 CSTEINMANNI_LOCUS2290461 (fragment)   Coregonus sp. balchen CAB1331594.1    
 hyaluronidase (fragment)   Crotalus adamanteus AEJ31976.1    
 hyaluronidase 1   Crotalus adamanteus AFJ49188.1    
 ORF (fragment)   Crotalus adamanteus AFJ49189.1    
 ORF (probable fragment)   Culex quinquefasciatus VERO BEACH AAR18444.1    
 hyaluronidase   Culicoides nubeculosus ACM40915.1    
 hyaluronidase / allergen (Cul n 2)   Culicoides nubeculosus ADJ67266.1    
 hyaluronidase / allergen (Cul o 2)   Culicoides obsoletus AGI16775.1    
 hyaluronidase (Hgs1;CsHGS1) (fragment)   Culicoides sonorensis AAU06477.1    
 hyaluronidase / endo-β-N-acetyl-hexosaminidase (CsHyal) 3.2.1.35 Cupiennius salei CUH74573.1 A0A0S4JYH2  
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