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Glycoside Hydrolase Family 56

Activities in FamilyChondroitinase / chondroitin hydrolase (EC 3.2.1.-);
Hyaluronidase / hyaluronoglucosaminidase (EC 3.2.1.35);
Mechanism Retaining e
3D Structure Status( β / α ) 7 barrel
Catalytic Nucleophile/Basecarbonyl oxygen of C-2 acetamido group of substrate
Catalytic Proton DonorGlu
External resourcesCAZypedia; PRINTS;
Statistics GenBank accession (532); Uniprot accession (63); PDB accession (6); 3D entries (3); cryst (0)
Summary
Download GH56 (372) Taxonomic display Structure (3) Characterized (29)
Eukaryota
Protein Name EC#OrganismGenBank UniprotPDB/3D
 hyaluronidase   Ampulex compressa ARK19841.1    
 venom hyaluronidase, partial (fragment)   Androctonus bicolor AIX87602.1    
 hyaluronidase   Androctonus crassicauda QQS80959.1    
 hyaluronidase (Hase-1)   Androctonus crassicauda UCR74866.1    
 hyaluronidase (HyaL)   Anoplius samariensis BAF93867.1    
 hyaluronidase 2   Apis cerana cerana AAK51798.2    
 hyaluronidase (HYA;Api m 2;Apim2;LOC406146) 3.2.1.35 Apis mellifera AAA27730.1
AAB45920.1
AAR78242.1
ABH87698.1
NP_001011619.1		 Q08169 1FCQ[A]
1FCU[A]
1FCV[A]
2J88[A]
 hyaluronidase   Apis mellifera carnica AFI40555.1    
 ORF   Ascaris suum ADY46439.1    
 Hyal1 (fragment)   Balaenoptera acutorostrata AEB78445.1    
 Hyal1 (fragment)   Balaenoptera omurai ADZ57824.1    
 BEMITA_ LOCUS 10475   Bemisia tabaci CAH0391897.1    
 LolHyaz (fragment)   Bichromomyia olmeca ANW11472.1    
 hyaluronidase   Bitis arietans ABI33944.1
ABI33945.1
ABI33946.1
ABI33947.1
ABI33948.1		    
 PH-20 hyaluronidase   Blarina brevicauda QKV49747.1    
 hyaluronidase, partial   Boiga irregularis AWX67683.1    
 Hyal2   Bos taurus AAL78385.1
AAI02043.1
AAX46601.1
AAX46657.1
ABG66994.1
NP_776772.1		    
 hyaluronidase (Spam1;PH-20;p80;bovHyal) 3.2.1.35 Bos taurus AAI10184.1
AAP55713.1
AAV41022.1		 Q2YDK3
Q5U7L5
Q7YS45		  
 HYAL1 (fragment)   Bos taurus AAX08792.1    
 HYAL1   Bos taurus AAX09052.1
AAI02474.1		    
 hyaluronidase   Bothrops atrox AUF71538.1    
 Hyaluronidase BmooHyal-2, partial   Bothrops moojeni ATU85543.1    
 Hyaluronidase BmooHyal-1, partial   Bothrops moojeni ATU85542.1    
 hyaluronidase (Hyase;BpHyase) (fragment)   Bothrops neuwiedi pauloensis ACN58137.1    
 hyaluronidase (Hyal;BvHyal) (partial) 3.2.1.35 Brachypelma vagans AFS33217.1 J9XYC6  
 MELIAE_ LOCUS 2367 (fragment)   Brassicogethes aeneus CAH0549075.1    
 BINO364_ LOCUS 2122   Brenthis ino CAH0715151.1    
 hyaluronidase PH-20, partial (fragment)   Bubalus bubalis bubalis ANC49763.1    
 hyaluronidase PH-20, partial (fragment)   Bubalus bubalis bubalis ANC49764.1    
 L3Y34_017400   Caenorhabditis briggsae isolate QX1410_ONT ULU04609.1    
 hyaluronidase / chondroitinase (Hyα-1;T22C8.2) 3.2.1.-
3.2.1.35		 Caenorhabditis elegans Bristol N2 CAA88874.1
BAG48551.1
NP_495830.1		 G5ECE8
Q22675		  
 7969   Cafeteria roenbergensis BVI (Cafroe1)      
 7720   Cafeteria roenbergensis BVI (Cafroe1)      
 PH-20   Callimico goeldii ACV60225.1    
 PH-20 (fragment)   Callimico goeldii ABD94489.1    
 hyaluronidase   Calliophis bivirgatus QSI83992.1    
 HYAL3-like protein (fragment)   Callithrix jacchus ABQ22669.1    
 ORF   Callorhinchus milii AFO99844.1    
 ORF   Callorhinchus milii AFO97416.1    
 ORF   Callorhinchus milii AFP02624.1    
 Hyal1 (fragment)   Camelus bactrianus AEB78448.1    
 sperm membrane protein (PH-20) (fragment)   Capra hircus ABI31753.1    
 hyaluronidase PH-20   Cavia porcellus CAA39768.1
AAB45922.1		    
 hyaluronidase   Cerastes cerastes ABI33938.1
ABI33939.1
ABI33940.1		    
 ORF (fragment)   Ceratonova shasta UKS51845.1    
 Hyal1 (fragment)   Ceratotherium simum AEB78449.1    
 PH-20   Cercopithecus mitis ACV60221.1    
 PH-20 (fragment)   Cercopithecus mitis ABD94493.1    
 CEUTPL_ LOCUS 8307 (fragment)   Ceutorhynchus assimilis CAG9767750.1    
 CEUTPL_ LOCUS 8310 (fragment)   Ceutorhynchus assimilis CAG9767753.1    
 CEUTPL_ LOCUS 8308 (fragment)   Ceutorhynchus assimilis CAG9767751.1    
 CEUTPL_ LOCUS 8309 (fragment)   Ceutorhynchus assimilis CAH1129628.1    
 venom protein (A4YE08CM1) (fragment)   Chelonus inanitus CBN72517.1    
 venom protein (A3YM10CM1) (fragment)   Chelonus inanitus CBN72518.1    
 venom protein (A2YM17CM2) (fragment)   Chelonus inanitus CBN72519.1    
 venom protein (A1YB14CM3) (fragment)   Chelonus inanitus CBN72520.1    
 CHILSU_ LOCUS 6564   Chilo suppressalis CAH0403295.1    
 CHIRRI_ LOCUS 55   Chironomus riparius CAG9797054.1    
 CHIRRI_ LOCUS 4158   Chironomus riparius CAG9801227.1    
 posterior head protein 20, partial (Ph-20)   Chlorocebus aethiops AEX20366.1    
 CINC_ LOCUS 8013   Chrysodeixis includens CAH0598162.1    
 395682   Ciona intestinalis (Cioin2)      
 378054   Ciona intestinalis (Cioin2)      
 posterior head protein 20, partial (Ph-20)   Colobus guereza AEX20365.1    
 Conohyal-Ad1 (fragment)   Conus adamsonii AFH78529.1    
 hyaluronidase (Conohyal-Cn1) 3.2.1.35 Conus consors IV AFH78528.1 I0CME7  
 venom-related protein conohyaluronidase, partial   Conus ebraeus UMA82668.1    
 conohyaluronidase   Conus ermineus AXL95480.1    
 conohyaluronidase, partial (fragment)   Conus ermineus AXL95522.1    
 conohyaluronidase, partial (fragment)   Conus ermineus AXL95351.1    
 conohyaluronidase, partial (fragment)   Conus magus QFQ61159.1    
 conohyaluronidase   Conus magus QFQ61160.1    
 conohyaluronidase   Conus magus QFQ61157.1    
 conohyaluronidase   Conus magus QFQ61158.1    
 ORF (fragment)   Coptotermes formosanus AGM32945.1    
 CSTEINMANNI_LOCUS2772392 (fragment)   Coregonus sp. balchen CAB1336289.1    
 CSTEINMANNI_LOCUS2772377   Coregonus sp. balchen CAB1336274.1    
 CSTEINMANNI_LOCUS3956487   Coregonus sp. balchen CAB1345787.1    
 CSTEINMANNI_LOCUS2290463   Coregonus sp. balchen CAB1331596.1    
 CSTEINMANNI_LOCUS352570   Coregonus sp. balchen CAB1314663.1    
 CSTEINMANNI_LOCUS4885995 (fragment)   Coregonus sp. balchen CAB1353427.1    
 CSTEINMANNI_LOCUS1271682   Coregonus sp. balchen CAB1323488.1    
 CSTEINMANNI_LOCUS4886719   Coregonus sp. balchen CAB1354151.1    
 CSTEINMANNI_LOCUS2290462   Coregonus sp. balchen CAB1331595.1    
 CSTEINMANNI_LOCUS1271681   Coregonus sp. balchen CAB1323487.1    
 CSTEINMANNI_LOCUS2048482   Coregonus sp. balchen CAB1329445.1    
 CSTEINMANNI_LOCUS2048494   Coregonus sp. balchen CAB1329457.1    
 CSTEINMANNI_LOCUS353316   Coregonus sp. balchen CAB1315409.1    
 CSTEINMANNI_LOCUS1271683   Coregonus sp. balchen CAB1323489.1    
 CSTEINMANNI_LOCUS4885994   Coregonus sp. balchen CAB1353426.1    
 CSTEINMANNI_LOCUS2290461 (fragment)   Coregonus sp. balchen CAB1331594.1    
 hyaluronidase (fragment)   Crotalus adamanteus AEJ31976.1    
 hyaluronidase 1   Crotalus adamanteus AFJ49188.1    
 ORF (fragment)   Crotalus adamanteus AFJ49189.1    
 ORF (probable fragment)   Culex quinquefasciatus VERO BEACH AAR18444.1    
 hyaluronidase   Culicoides nubeculosus ACM40915.1    
 hyaluronidase / allergen (Cul n 2)   Culicoides nubeculosus ADJ67266.1    
 hyaluronidase / allergen (Cul o 2)   Culicoides obsoletus AGI16775.1    
 hyaluronidase (Hgs1;CsHGS1) (fragment)   Culicoides sonorensis AAU06477.1    
 hyaluronidase / endo-β-N-acetyl-hexosaminidase (CsHyal) 3.2.1.35 Cupiennius salei CUH74573.1 A0A0S4JYH2  
 Unknown (protein for MGC:136804)   Danio rerio AAI14278.1    
 DKEY-42I9.14-001 (Hyal2)   Danio rerio CAK11026.1    
 DKEY-72L14.3-001   Danio rerio CAK11158.1    
 Unknown (protein for MGC:158626)   Danio rerio AAI29331.1    
 Unknown (protein for MGC:158668)   Danio rerio AAI29368.1
AAI52242.1		    
 DKEY-20D21.13-001   Danio rerio CAP19590.1    
 Hyal1 (fragment)   Delphinapterus leucas ADZ57829.1    
 Hyal1 (fragment)   Delphinus capensis ADZ57818.1    
 DIABBA_ LOCUS 7349 (fragment)   Diabrotica balteata CAG9833995.1    
 hyaluronidase (Dol m 2) 3.2.1.35 Dolichovespula maculata AAA68279.1
AAR78243.1
ABH87699.1
CAJ98548.1		 P49371  
 venom polypeptide precursor   Doratifera vulnerans QTY40918.1    
 hyaluronidase (fragment)   Echis carinatus sochureki ABI33949.1    
 hyaluronidase (fragment)   Echis carinatus sochureki ABI33950.1    
 hyaluronidase (fragment)   Echis ocellatus ABI33937.1    
 hyaluronidase (fragment)   Echis pyramidum leakeyi ABI33941.1    
 hyaluronidase (fragment)   Echis pyramidum leakeyi ABI33942.1
ABI33943.1		    
 Hyal1 (fragment)   Elaphurus davidianus ADZ57827.1    
 hyaluronoglucosaminidase 1 (Has1)   Eospalax fontanierii baileyi ANJ16065.1    
 sperm adhesion molecule 1 (Spam1) (fragment)   Equus caballus AET34452.1    
 hyaluronoglucosaminidase 1, partial (Hyal1) (fragment)   Erinaceus europaeus AEA40788.1    
 PH-20   Erythrocebus patas ACV60222.1    
 PH-20 (fragment)   Erythrocebus patas ABD94492.1    
 v-hyaluronidase-Lio1 (fragment)   Erythrolamprus poecilogyrus ABU68570.1    
 hyaluronidase   Eulalia sp. (in: segmented worms) WWC50937.1    
 venom hyaluronidase (Hya1)   Eumenes pomiformis ACZ37402.1    
 hyaluronidase-Ginf1 (fragment)   Gerrhonotus infernalis ADK39240.1    
 hyaluronoglucosaminidase   Gloydius intermedius ALB06111.1    
 GFP_L7_0750 (possible fragment)   Glyptapanteles flavicoxis ACE75133.1    
 GFP_L7_0760 (possible fragment)   Glyptapanteles flavicoxis ACE75134.1    
 GFP_L7_0770 (fragment)   Glyptapanteles flavicoxis ACE75135.1    
 GIP_L1_00710   Glyptapanteles indiensis ABK57057.1    
 GMON_LOCUS5768 (fragment)   Gordionus sp. m RMFG-2023 CAK9298537.1    
 GMON_LOCUS5103   Gordionus sp. m RMFG-2023 CAK9297611.1    
 GMON_LOCUS8880   Gordionus sp. m RMFG-2023 CAK9303253.1    
 GMON_LOCUS5826   Gordionus sp. m RMFG-2023 CAK9298627.1    
 GMON_LOCUS8877   Gordionus sp. m RMFG-2023 CAK9303248.1    
 GMON_LOCUS8879 (fragment)   Gordionus sp. m RMFG-2023 CAK9303251.1    
 GMON_LOCUS5769 (fragment)   Gordionus sp. m RMFG-2023 CAK9298538.1    
 GMON_LOCUS8876   Gordionus sp. m RMFG-2023 CAK9303247.1    
 GMON_LOCUS10003   Gordionus sp. m RMFG-2023 CAK9304945.1    
 posterior head protein 20, partial (Ph-20)   Gorilla gorilla AEX20363.1    
 Hyal1 (fragment)   Grampus griseus ADZ57819.1    
 hyaluronidase toxin (fragment)   Heloderma suspectum cinctum ACE95063.1    
 venom toxin   Hemiscorpius lepturus API81375.1    
 hyaluronidase, partial (fragment)   Hemiscorpius lepturus QQS80960.1    
 hyaluronidase (Hase1)   Hemiscorpius lepturus UCR74869.1    
 hyaluronidase (Hase2)   Hemiscorpius lepturus UCR74868.1    
 hyaluronidase (Hase3)   Hemiscorpius lepturus UCR74867.1    
 HERILL_LOCUS10861   Hermetia illucens CAD7088215.1    
 Hyal1 (fragment)   Hippopotamus amphibius AEB78446.1    
 hyaluronidase (SPAM1) (sperm) 3.2.1.35 Homo sapiens AAH26163.1
AAC16482.1
AAC60607.2
AAQ96882.1
AAX13985.1
AAX13986.1
AAX13987.1
AAX13988.1
AAY03322.1
ABJ38775.1
ACJ13740.1
ACM84481.1
BAF84918.1
CAA59086.1
NP_003108.2
NP_694859.1		 P38567
Q5D1J4
Q8TC30		  
 chondroitin sulfate hydrolase / hyaluronidase 3 (HYAL3;LUCA-3;LUCA14.2) 3.2.1.-
3.2.1.35		 Homo sapiens AAC70915.1
AAC02729.1
AAD04257.1
AAH05896.1
AAH12892.1
AAM60775.1
AAM60776.1
AAM60777.1
AAM60778.1
ABA67787.1
BAG36242.1
NP_003540.2		 B2R8P5
O43820
O60540
Q8NFK2
Q8NFK3
Q8NFK4
Q96E56
Q9BRW9		  
 chondroitin sulfate hydrolase / hyaluronidase 1 (HYAL1;Hyal-1;LUCA1;LUCA-1) 3.2.1.-
3.2.1.35		 Homo sapiens AAD09137.2
AAB67046.1
AAC02730.1
AAD04190.1
AAD53277.1
AAE60457.1
AAE60459.1
AAH25774.1
AAH35695.1
AAM60770.1
AAM60771.1
AAM60772.1
AAM60773.1
AAM60774.1
ABJ38770.1
ABJ38772.1
ACK03148.1
ALQ33600.1
ALQ33601.1
BAF83505.1
BAG53447.1
CAC60225.1
CAG46731.1
NP_009296.1
NP_149349.2
NP_695013.1
NP_695014.1
NP_695015.1
NP_695016.1
NP_695017.1
NP_695018.1		 A0A024R2X3
A0A0S2Z3Q0
A8K451
B3KUI5
Q12794
Q6FH23		 2PE4[A]
 chondroitin sulfate hydrolase / hyaluronidase 2 (Hyal2;LUCA2;LUCA-2) 3.2.1.-
3.2.1.35		 Homo sapiens AAH00692.1
AAB67045.1
AAC28656.1
AAC62823.1
ABA82488.1
ACK03149.1
BAG52554.1
BAG54602.1
CAA03924.1
NP_003764.3
NP_149348.2		 B3KRZ2
Q12891		  
 chondroitin sulfate hydrolase / hyaluronoglucosaminidase 4 / hyaluronidase 4 (Hyal4;HYAL4;HYAL-4) 3.2.1.- Homo sapiens AAC98883.1
AAD43186.1
AAI04789.1
AAI04791.1
ACM81328.1
ACM82678.1
ACV97636.1
BAI49593.1
NP_036401.1
QIJ31491.1
QIJ31492.1		 A0A6G7MAH8
A0A6G7MAX3
D0VXG1
Q2M3T9
Q9UL99		  
 ENSP00000194108 (fragment)   Homo sapiens EAW83601.1
XP_069745.1
XP_069745.2		    
 ORF (fragment)   Hottentotta judaicus ADY39497.1    
 PH-20   Hylobates lar ACV60218.1    
 PH-20 (fragment)   Hylobates lar ABD94497.1    
 ORF (Hyal2)   Ictalurus punctatus AHH39098.1    
 ORF (Hyal1)   Ictalurus punctatus AHH42852.1    
 IPOD504_ LOCUS 10363   Iphiclides podalirius CAH2057920.1    
 IPOD504_ LOCUS 10363   Iphiclides podalirius CAH2057922.1    
 Hyal1 (fragment)   Kogia sima ADZ57825.1    
 venom protein   Lampona murina WBW70090.1    
 venom protein   Lampona murina WBW70091.1    
 PH-20 (fragment)   Lemur catta ABD94483.1    
 PH-20   Leontocebus fuscicollis ACV60227.1    
 PH-20 (fragment)   Leontocebus fuscicollis ABD94488.1    
 LSINAPIS_LOCUS6006   Leptidea sinapis VUY93948.1    
 LSINAPIS_LOCUS5075 (fragment)   Leptidea sinapis VUY92740.1    
 venom hyaluronidase   Lethocerus distinctifemur ATU82477.1    
 Hyal1 (fragment)   Lipotes vexillifer ADZ57817.1    
 PH-20   Lophocebus aterrimus ACV60228.1    
 PH-20 (fragment)   Lophocebus aterrimus ABD94495.1    
 hyaluronidase 3.2.1.35 Loxosceles intermedia AGH25912.1 R4J7Z9  
 hyaluronidase (HyaL)   Lutzomyia longipalpis AAD32195.1    
 ORF (fragment)   Lycosa tarantula QNF22879.1    
 ORF (fragment)   Lycosa tarantula QNF22877.1    
 hyaluronidase   Macaca fascicularis BAE01371.1
ABD94494.1		    
 PH-20   Macaca fascicularis ACV60220.1    
 hyaluronidase-2 (Hyal2)   Macaca mulatta AFI35008.1
AFE69962.1		    
 hyaluronidase, partial (fragment)   Macrovipera lebetina AHJ80887.1    
 Hyal1 (fragment)   Manis pentadactyla AEB78450.1    
 venom protein U-MPTX.13-Mc23, partial (fragment)   Megalopyge crispata WJJ70432.1    
 venom protein U-MPTX.13-Mc36, partial (fragment)   Megalopyge crispata WJJ70431.1    
 venom protein U-MPTX.13-30 (fragment)   Megalopyge opercularis WJJ70397.1    
 hyaluronidase   Mesobuthus eupeus WWA73958.1    
 hyaluronidase (Hya1;BmHYA1) (partial) 3.2.1.35 Mesobuthus martensii ACY69673.1
AHA36327.1		 D1MBU1
P86100		  
 SPAM1   Mesocricetus auratus ADB94578.1    
 hyaluronidase 5 (Hyal5)   Mesocricetus auratus ADB94579.1    
 hyaluronoglucosaminidase 1 (Hyal1) (fragment)   Mesoplodon densirostris ADZ53990.1    
 hyaluronidase   Meteorus pulchricornis BAL70300.1    
 PH-20 (fragment)   Microcebus murinus ABD94486.1    
 Hyal1 (fragment)   Moschus moschiferus ADZ57828.1    
 hyaluronidase 3 (Hyal3)   Mus musculus AAH18457.1
AAL06145.1
CAP19324.1
NP_821139.1		    
 hyaluronidase (Spam1;PH-20;Spam) 3.2.1.35 Mus musculus AAA76603.1
AAP49832.1
AAX13989.1
AAX13990.1
BAB24161.1
BAC55070.1
NP_033267.1		 P48794
Q5D1J0		  
 hyaluronoglucosaminidase 1 (Hyal1;Hyal-1) 3.2.1.35 Mus musculus AAC15949.1
AAH21636.1
AAL17822.1
BAC26206.1
CAP19319.1
CAP19320.1
CAP19321.1
CAP19322.1
NP_032343.2		 O70229
Q8CE62
Q8VDK0
Q91ZJ9		  
 hyaluronidase (Hyal-5;Hyal5)   Mus musculus BAC55071.1
AAI45993.1
AAI45995.1
BAB30602.2
BAB30602.3
BAE21628.1		    
 hyaluronidase 2 (Hyal2;Hyal-2) 3.2.1.35 Mus musculus AAK28482.1
AAH60047.1
AAK28481.1
AAL17823.1
AAO49503.1
BAC39851.1
BAC40583.1
BAE21913.1
CAA03888.1
CAA03889.1
NP_034619.1		 O35632
Q3UZE4
Q542F2
Q8C309		  
 4932701A20 or 4930503G08   Mus musculus C57BL/6J BAB30316.1
BAC26578.1		    
 chondroitin sulfate hydrolase (Hyal4;mHYAL4) 3.2.1.- Mus musculus C57BL/6J BAB29454.1
AAI25403.1
AAI32097.1
BAM45641.1		 L0MX76
Q05A56
Q9D660		  
 ORF (fragment)   Mustela putorius furo AER99997.1    
 hyaluronoglucosaminidase 1 (fragment)   Mustela putorius furo AER99995.1    
 hyaluronoglucosaminidase 2   Mustela putorius furo AER99996.1    
 ORF (fragment)   Mustela putorius furo AES10768.1    
 ORF (fragment)   Mustela putorius furo AES10769.1    
 MAR_004820 (fragment)   Mya arenaria isolate MELC-2E11 WAR14715.1    
 MAR_004820 (fragment)   Mya arenaria isolate MELC-2E11 WAR14715.1    
 MAR_004800 (fragment)   Mya arenaria isolate MELC-2E11 WAR14695.1    
 posterior head protein 20, partial (Ph-20)   Nasalis larvatus AEX20364.1    
 Hyal1 (fragment)   Neophocaena phocaenoides ADZ57816.1    
 NTJ_04684   Nesidiocoris tenuis Japan BES91875.1    
 NEZAVI_ LOCUS 12776 (fragment)   Nezara viridula CAH1404348.1    
 NEZAVI_ LOCUS 9411   Nezara viridula CAH1400109.1    
 Q1695_012661   Nippostrongylus brasiliensis MIMR WKX96390.1    
 Q1695_013453 (fragment)   Nippostrongylus brasiliensis MIMR WKX97786.1    
 Q1695_013157 (fragment)   Nippostrongylus brasiliensis MIMR WKX97271.1    
 Q1695_013452 (fragment)   Nippostrongylus brasiliensis MIMR WKX97783.1    
 Q1695_004957   Nippostrongylus brasiliensis MIMR WKY03612.1    
 hyaluronidase (fragment)   Nyssomyia intermedia AFP99265.1    
 OCTVUL_1B014278   Octopus vulgaris CAI9731055.1    
 OCTVUL_1B007469   Octopus vulgaris CAI9731056.1    
 OCTVUL_1B007469 (fragment)   Octopus vulgaris CAI9731057.1    
 hyaluronidase (Hyal_oM) (fragment)   Odontomachus monticola BBF97841.1    
 hyaluronidase   Orancistrocerus drewseni ACD61711.1    
 hyauronidase, partial   Ornithorhynchus anatinus AFV61285.1    
 hyaluronidase 2 (Hyal2)   Oryctolagus cuniculus AAT35556.1    
 hyaluronidase (Hyal1;Spam1;PH20)   Oryctolagus cuniculus AAA88913.1    
 PH-20 (fragment)   Otolemur crassicaudatus ABD94481.1    
 Hyal2   Ovis aries AAL78386.1    
 hyaluronidase 2 / hyaluronoglucosaminidase 2 (Hyal2)   Ovis aries AFP89957.1    
 hyaluronidase   Ovophis okinavensis BAN89413.1
BAN89414.1		    
 ORF   Pan troglodytes BAK64019.1    
 ORF   Pan troglodytes BAK62198.1    
 ORF (fragment)   Pan troglodytes BAK62386.1    
 PH-20 (fragment)   Pan troglodytes ABD94498.1    
 PH-20   Papio hamadryas ACV60219.1    
 PH-20 (fragment)   Papio hamadryas ABD94496.1    
 PECUL_23A031060   Pelobates cultripes CAH2278175.1    
 PECUL_23A043003 (fragment)   Pelobates cultripes CAH2275809.1    
 PECUL_23A049834   Pelobates cultripes CAH2312554.1    
 PECUL_23A049834   Pelobates cultripes CAH2312554.1    
 PECUL_23A049834   Pelobates cultripes CAH2312554.1    
 PH-20 (fragment)   Perodicticus potto ABD94482.1    
 PHAECO_ LOCUS 2625 (fragment)   Phaedon cochleariae CAH1118722.1    
 ORF   Phallusia mammillata CAB3254999.1    
 salivary hyaluronidase   Phlebotomus arabicus ACS93505.1    
 salivary hyaluronidase (fragment)   Phlebotomus orientalis Addis Zemen AGT96452.1    
 salivary hyaluronidase (fragment)   Phlebotomus orientalis Melka Werer AGT96475.1    
 hyaluronidase   Phlebotomus perniciosus ALL27024.1    
 ORF (fragment)   Phlebotomus tobbi AEK98519.1    
 PHYEVI_ LOCUS 4559   Phyllotreta striolata CAG9858168.1    
 ORF (fragment)   Pipa carvalhoi AEQ16640.1    
 MPIPNATIZW_LOCUS4184   Pipistrellus nathusii CAK6435878.1    
 MPIPNATIZW_LOCUS2586   Pipistrellus nathusii CAK6434280.1    
 MPIPNATIZW_LOCUS4183   Pipistrellus nathusii CAK6435877.1    
 MPIPNATIZW_LOCUS4186   Pipistrellus nathusii CAK6435880.1    
 MPIPNATIZW_LOCUS2587 (fragment)   Pipistrellus nathusii CAK6434281.1    
 Hyal1 (fragment)   Platanista gangetica ADZ57826.1    
 venom hyaluronidase 1   Platymeris rhadamanthus QHB21608.1    
 PODLI_1B039194   Podarcis lilfordi CAI5767050.1    
 PODLI_1B040523   Podarcis lilfordi CAI5767047.1    
 PODLI_1B039257   Podarcis lilfordi CAI5767867.1    
 PODLI_1B002495   Podarcis lilfordi CAI5767111.1    
 PODLI_1B036710   Podarcis lilfordi CAI5767049.1    
 PODLI_1B000770   Podarcis lilfordi CAI5785342.1    
 PODLI_1B033037   Podarcis lilfordi CAI5785343.1    
 hyaluronidase (Pol a 2)   Polistes annularis AAD52616.1
AAR78239.1
ABH87695.1		    
 Hyal / Pp-Hyal   Polybia paulista ADL09135.1    
 DKFZp469J0738   Pongo abelii CAH89756.1    
 PH-20 (fragment)   Propithecus verreauxi ABD94485.1    
 hyaluronidase, partial (fragment)   Protobothrops flavoviridis BAN89379.1
BAP39986.1		    
 hyaluronidase   Pseudocerastes urarachnoides UMW88234.1    
 hyaluronidase (fragment)   Pseudocerastes urarachnoides UMW88233.1    
 hyaluronidase   Pseudonaja textilis WVG99541.1    
 PSYICH_ LOCUS 15505   Psylliodes chrysocephala CAH1115357.1    
 hyaluronidase 3.2.1.35 Pterois antennata BAM66299.1 K7ZPU7  
 hyaluronidase 3.2.1.35 Pterois volitans BAM66300.1 K7ZMF5  
 PH-20   Pygathrix nemaeus ACV60224.1    
 PH-20 (fragment)   Pygathrix nemaeus ABD94490.1    
 MRATA1EN1_LOCUS9565   Rangifer tarandus platyrhyncus CAI9160603.1    
 MRATA1EN1_LOCUS9566   Rangifer tarandus platyrhyncus CAI9160604.1    
 MRATA1EN1_LOCUS9567   Rangifer tarandus platyrhyncus CAI9160605.1    
 MRATA1EN1_LOCUS8640   Rangifer tarandus platyrhyncus CAI9159678.1    
 MRATA1EN1_LOCUS8638   Rangifer tarandus platyrhyncus CAI9159676.1    
 hyaluronidase 1 (Hyal1)   Rattus norvegicus BAD14368.1    
 hyaluronidase 4 (Hyal4) (fragment)   Rattus norvegicus BAD14369.1    
 hyaluronidase 3 (Hyal3)   Rattus norvegicus BAD14370.1
AAH97259.1		    
 Unknown (protein for MGC:94293)   Rattus norvegicus AAH82016.1    
 Unknown (protein for MGC:108951)   Rattus norvegicus AAH91219.1    
 2B1 antigen   Rattus norvegicus AAH81748.1
CAA62016.1
EDM15179.1
NP_446419.1		    
 hyaluronidase 2 (Hyal2) 3.2.1.35 Rattus norvegicus AAD01980.1
AAH62410.1
AAO49504.1
NP_742037.1		 Q80ZC7
Q9Z2Q3		  
 venom hyaluronidase   Rhynchium brunneum ACC93934.1    
 posterior head protein 20, partial (Ph-20)   Saguinus oedipus AEX20362.1    
 PH-20   Saimiri sciureus ACV60226.1    
 PH-20 (fragment)   Saimiri sciureus ABD94487.1    
 hyaluronidase-2 (Hyal2)   Salmo salar ACI32917.1    
 hyaluronidase-2 (Hyal2)   Salmo salar ACI33835.1    
 hyaluronidase-2 (Hyal2)   Salmo salar ACI33947.1
ACN10738.1		    
 SMAX5B_011659   Scophthalmus maximus AWP07919.1    
 SMAX5B_011659   Scophthalmus maximus AWP07919.1    
 SMAX5B_011659   Scophthalmus maximus AWP07919.1    
 SMAX5B_002348   Scophthalmus maximus AWP02649.1    
 SMAX5B_002348   Scophthalmus maximus AWP02649.1    
 SMAX5B_002219   Scophthalmus maximus AWP02765.1    
 SMAX5B_007737   Scophthalmus maximus AWO98026.1    
 SMAX5B_002682   Scophthalmus maximus AWP09387.1    
 SMAX5B_002618   Scophthalmus maximus AWP09369.1    
 PH-20   Semnopithecus entellus ACV60223.1    
 PH-20 (fragment)   Semnopithecus entellus ABD94491.1    
 salivary hyaluronidase, partial (fragment)   Sergentomyia schwetzi QHO60719.1    
 salivary hyaluronidase, partial (fragment)   Sergentomyia schwetzi QHO60720.1    
 salivary hyaluronidase   Sergentomyia schwetzi QHO60718.1    
 salivary hyaluronidase   Simulium guianense AEB96520.1    
 Hyal1 (fragment)   Sousa chinensis ADZ57823.1    
 SPEXI_ LOCUS 6935   Spodoptera exigua CAH0697278.1    
 SPLIT_ LOCUS 7052   Spodoptera littoralis CAH1641696.1    
 Hyal1 (fragment)   Stenella attenuata ADZ57820.1    
 Hyal1 (fragment)   Stenella coeruleoalba ADZ57822.1    
 SRAE_2000096800   Strongyloides ratti ED321 Heterogonic CEF66298.1    
 SRAE_X000016600   Strongyloides ratti ED321 Heterogonic CEF70836.1    
 SRAE_2000275100   Strongyloides ratti ED321 Heterogonic CEF68093.1    
 SRAE_X000016700   Strongyloides ratti ED321 Heterogonic CEF70837.1    
 SRAE_X000016500 (fragment)   Strongyloides ratti ED321 Heterogonic CEF70835.1    
 sperm adhesion molecule 1 (SPAM-1)   Sus scrofa CAC83973.2    
 hyaluronoglucosaminidase 3   Sus scrofa AAR91601.1
AAR33037.1		    
 hyaluronoglucosaminidase 1   Sus scrofa AAR91599.1
ACH56985.1
ACH56986.1		    
 hyaluronidase 2 (Hyal2)   Sus scrofa AAR91600.1
ACH56987.1		    
 Hyal1 (fragment)   Sus scrofa AEB78447.1    
 hyaluronidase PH-20, partial   Symphalangus syndactylus AEH95769.1    
 hyaluronidase (HYA1;SfHya1) 3.2.1.35 Synanceia horrida AAO74499.1 Q801Z8  
 hyaluronidase (Hya1;SvHya1;Sv-Hya)   Synanceia verrucosa BAJ54082.1    
 hyaluronidase TY / allergen (Tab y 2)   Tabanus yao ADM18346.1    
 TDIB3V08_LOCUS196   Timema douglasi CAD7193756.1    
 TSIB3V08_LOCUS1818 (fragment)   Timema shepardi CAD7257560.1    
 ORF   Tityus cisandinus WDU65909.1    
 ORF   Tityus melici WLF82748.1
WLF82749.1
WLF82750.1
WLF82751.1
WLF82752.1
WLF82753.1		    
 ORF   Tityus serrulatus QPD99039.2
QPD99039.1		    
 ORF   Tityus serrulatus QPD99040.2
QPD99040.1		    
 hyaluronidase 2 (Hyal-2;TsHyal-2) 3.2.1.35 Tityus serrulatus AHF72516.1 W0HFN9  
 hyaluronidase 1 (Hya; Hyal-1; TsHyal-1) 3.2.1.35 Tityus serrulatus AHF72517.1 P85841  
 Hyal1 (fragment)   Tursiops aduncus ADZ57821.1    
 Hyal1 (fragment)   Tursiops truncatus AEB78444.1    
 PH-20 (fragment)   Varecia variegata ABD94484.1    
 hyaluronidase (Vesp ma 2) 3.2.1.35 Vespa magnifica CBY83816.1 P86875  
 hyaluronidase type B (HAase b; VesT2b)   Vespa tropica UYX46121.1    
 hyaluronidase (Ves g 2a)   Vespula germanica CAL59818.1    
 Ves g 2b (fragment)   Vespula germanica CAL59819.1    
 hyaluronidase b (Hya_B;Ves v 2b)   Vespula vulgaris AAX14718.1
CAI77218.1		    
 hyaluronidase (Ves v 2 allergen) 3.2.1.35 Vespula vulgaris AAB48073.1
AAR78244.1
ABH87700.1		 P49370 2ATM[A]
 hyaluronidase   Vipera anatolica senliki QHR82765.1    
 fPH-20 protein   Vulpes vulpes AAB86588.1    
 hyaluronidase XKH1 3.2.1.35 Xenopus laevis AAL67683.1
AAI69767.1
AAI69795.1		 B7ZQD4
Q8UVT7		  
 lysosomal hyaluronidase (Hyal2) 3.2.1.35 Xenopus laevis AAL55823.1 Q8UVY6  
 ORF   Xenopus laevis AAH47252.1    
 hyaluronidase (embryonic) XEH1 3.2.1.35 Xenopus laevis AAD28281.1 Q9W6K5  
 Unknown (protein for MGC:121214)   Xenopus tropicalis AAI35205.1    
 Unknown (protein for MGC:184955)   Xenopus tropicalis AAI58240.1    
 XNOV1_A013835   Xyrichtys novacula CAJ1056044.1    
 XNOV1_A023686   Xyrichtys novacula CAJ1056045.1    
 XNOV1_A010219   Xyrichtys novacula CAJ1056046.1    
 XNOV1_A041606   Xyrichtys novacula CAJ1055492.1    
 XNOV1_A032864   Xyrichtys novacula CAJ1055469.1    
 XNOV1_A010277   Xyrichtys novacula CAJ1056047.1    
 XNOV1_A006631   Xyrichtys novacula CAJ1050702.1    
 XNOV1_A021649   Xyrichtys novacula CAJ1054934.1    
 XNOV1_A017468   Xyrichtys novacula CAJ1050570.1    
 XNOV1_A037184   Xyrichtys novacula CAJ1050569.1    
 XNOV1_A037184   Xyrichtys novacula CAJ1050569.1    
Viruses
Protein Name EC#OrganismGenBank UniprotPDB/3D
 ORF   Bracoviriform congregatae CCQ71107.1    
 GIP_L8_0010 (fragment)   Glyptapanteles indiensis bracovirus ACE75387.1    
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