Activities in Sub Family	[retaining] α-1,4-glucan branching enzyme (EC 2.4.1.18);
Mechanism	Retaining a
Clan	GH-H
3D Structure Status	( β / α ) 8 barrel
Catalytic Nucleophile/Base	Asp (experimental)
Catalytic Proton Donor	Glu (experimental)
Note	New: many members have been assigned to subfamilies as described by Stam et al. (2006) Protein Eng Des Sel. 19, 555-562 (PMID: 17085431)
External resources	CAZypedia; EBI Protein of the Month; HOMSTRAD; PDB Molecule of the Month; PRINTS;
Commercial Enzyme Provider(s)	MEGAZYME; PROZOMIX;
Statistics	GenBank accession (3); Uniprot accession (32); PDB accession (26); 3D entries (6); cryst (0)

Bacteria
Protein Name	EC#	Organism	GenBank	Uniprot	PDB/3D Carbohydrate Ligands Resolution (Å)	Subf
α-1,4-glucan branching enzyme (BE3;cceBE3;cce_1806)	2.4.1.18	Crocosphaera subtropica ATCC 51142	ACB51156.1	B1WZK4	7XSY[A,B,C,D] 2.50	9
α-1,4-glucan branching enzyme (GlgB;BE1;cceBE1;cce_2248)	2.4.1.18	Crocosphaera subtropica ATCC 51142	ACB51598.1	B1WPM8	5GQU[A] 1.85 5GQV[A] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 3.00 5GQW[A] 1.80 5GQX[A] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 2.30 5GQY[A] α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 2.00 5GQZ[A] 1.85 5GR0[A] 1.95 5GR1[A] α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp 2.50 5GR2[A] 1.95 5GR3[A] 2.60 5GR4[A] α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-D-1-deoxy-Glcp 2.00 5GR5[A] 2.40 5GR6[A] 2.00 6KLF[A] 2.50	9
branching-enzyme (EcBE;GlgB;EcE24377A_3911)	2.4.1.18	Escherichia coli O139:H28 str. E24377A	ABV17243.1	A7ZSW5 P07762	5E6Y[A,B,C,D] a-cyclodextrin 2.60 5E6Z[A,B,C,D] b-cyclodextrin 1.88 5E70[A,B,C,D] 2.33	9
branching-enzyme (GlgB;EcoBE;b3432)	2.4.1.18	Escherichia coli str. K-12 substr. MG1655	AAA23872.1 AAA58230.1 AAC76457.1 ABX94236.1 AIZ89568.1 ALI38888.1 AMC96315.1 AUG18135.1 AVI53591.1 AYG17626.1 CAP11381.1 NP_417890.1	P07762	1M7X[A] 2.30 3O7Y[A,B,C,D] α-D-Glcp-(1-4)-D-1-deoxy-Glcp α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-β-D-Glcp 2.41 3O7Z[A,B,C,D] α-D-Glcp-(1-4)-α-D-Glcp β-D-Glcp 2.55 4LPC[A,B,C,D] α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-β-D-Glcp α-D-Glcp-(1-4)-β-D-Glcp β-D-Glcp 2.39 4LQ1[A,B,C,D] α-D-Glcp-(1-4)-α-D-Glcp β-D-Glcp 2.55 8SDB[A,B,C,D] α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp-(1-4)-α-D-Glcp 3.00	9
branching-enzyme (GlgB;Rv1326c)	2.4.1.18	Mycobacterium tuberculosis H37Rv	CAA98090.1 AFN49233.1 CCP44084.1 NP_215842.1	P9WN45 Q10625	3K1D[A] 2.33	9
branching enzyme (GlgB;RoBE)	2.4.1.18	Rhodothermus marinus JCM9785	BAB69858.1 AAR45540.1	Q93HU3	6JOY[A] 2.39	9
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