Carbohydrate-Binding Module Family 4

Activities in FamilyModules of approx. 150 residues found in bacterial enzymes. Binding of these modules has been demonstrated with xylan, β-1,3-glucan, β-1,3-1,4-glucan, β-1,6-glucan and amorphous cellulose but not with crystalline cellulose.
3D Structure Statusβ-sandwich
NotePreviously known as cellulose-binding domain family IV (CBD IV).
External resourcesCAZypedia; HOMSTRAD;
Statistics GenBank accession (526); Uniprot accession (125); PDB accession (18); 3D entries (10); cryst (4)
All (707) Archaea (2) Bacteria (697) unclassified (8) Structure (10 - 4 cryst) Characterized (53)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D
 CEH05_15895   Halobacillus halophilus HL2HP6 ASF40552.1    
 HM131_13645   Halobacillus mangrovi KTB 131 ARI77828.1    
 HM131_13645   Halobacillus mangrovi KTB 131 ARI77828.1    
 HM131_13645   Halobacillus mangrovi KTB 131 ARI77828.1    
 Halha_0267   Halobacteroides halobius DSM 5150 AGB40278.1    
 Halha_0267   Halobacteroides halobius DSM 5150 AGB40278.1    
 Halha_0268   Halobacteroides halobius DSM 5150 AGB40279.1    
 Halha_0268   Halobacteroides halobius DSM 5150 AGB40279.1    
 Hore_04240   Halothermothrix orenii H 168 ACL69182.1 B8D1V5  
 Hore_04240   Halothermothrix orenii H 168 ACL69182.1 B8D1V5  
 SD1D_1781 (CelK)   Herbinix luporum SD1D CUH93326.1    
 HVS_10850 (Celk2)   Herbivorax saccincola GGR1 AUG58065.1    
 HVS_07950 (Celk1)   Herbivorax saccincola GGR1 AUG57501.1    
 HVS_07950 (Celk1)   Herbivorax saccincola GGR1 AUG57501.1    
 HVS_07950 (Celk1)   Herbivorax saccincola GGR1 AUG57501.1    
 HVS_11170 (Celk3)   Herbivorax saccincola GGR1 AUG58129.1    
 HVS_10860   Herbivorax saccincola GGR1 AUG58067.1    
 Haur_1418   Herpetosiphon aurantiacus DSM 785 ABX04062.1 A9B399  
 Haur_1418   Herpetosiphon aurantiacus DSM 785 ABX04062.1 A9B399  
 IMCC20628_03694   Hoeflea sp. IMCC20628 AKI02378.1    
 Hsw_2363   Hymenobacter swuensis DY53 AHJ97958.1    
 I598_0211 (CenC)   Isoptericola dokdonensis DS-3 ANC29802.1    
 I598_0211 (CenC)   Isoptericola dokdonensis DS-3 ANC29802.1    
 JMA_31360   Jeotgalibacillus malaysiensis D5 AJD92453.1    
 JMA_31360   Jeotgalibacillus malaysiensis D5 AJD92453.1    
 JMA_31360   Jeotgalibacillus malaysiensis D5 AJD92453.1    
 Krad_3823   Kineococcus radiotolerans SRS30216 = ATCC BAA-149 ABS05286.1 A6WEP7  
 AM394_02270 (fragment)   Klebsiella oxytoca AR_0147 ARB20137.1    
 KPK_A0121 (CelK)   Klebsiella pneumoniae 342 ACI12123.1 B5RK45  
 Cphy_3388   Lachnoclostridium phytofermentans ISDg ABX43741.1 A9KTB1  
 Cphy_3388   Lachnoclostridium phytofermentans ISDg ABX43741.1 A9KTB1  
 Cphy_3388   Lachnoclostridium phytofermentans ISDg ABX43741.1 A9KTB1  
 Cphy_3388   Lachnoclostridium phytofermentans ISDg ABX43741.1 A9KTB1  
 Verru16b_01834   Lacunisphaera limnophila IG16b AOS44766.1    
 BBK82_39340   Lentzea guizhouensis DHS C013 ANZ41141.1    
 LPB138_07660   Lutibacter sp. LPB0138 AOW20562.1    
 LPB138_07660   Lutibacter sp. LPB0138 AOW20562.1    
 LPB138_07660   Lutibacter sp. LPB0138 AOW20562.1    
 MROS_0752   Melioribacter roseus P3M-2 AFN73994.1    
 xylanase (Xyl2091;MROS_2091) 3.2.1.8 Melioribacter roseus P3M-2 AFN75321.1    
 MEBOL_001951   Melittangium boletus DSM 14713 ATB28503.1    
 MEBOL_006774   Melittangium boletus DSM 14713 ATB33283.1    
 MEBOL_003465   Melittangium boletus DSM 14713 ATB30010.1    
 MEBOL_001952   Melittangium boletus DSM 14713 ATB28504.1    
 BO218_05495   Microbacterium paludicola CC3 APF33713.1    
 BO218_05495   Microbacterium paludicola CC3 APF33713.1    
 Micau_2188   Micromonospora aurantiaca ATCC 27029 ADL45733.1 D9T4M2  
 GA0070611_0362   Micromonospora auratinigra DSM 44815 SBT37878.1    
 GA0070612_5046   Micromonospora chokoriensis DSM 45160 SCF22474.1    
 GA0070612_5445   Micromonospora chokoriensis DSM 45160 SCF25646.1    
 GA0070607_0553   Micromonospora coriariae DSM 44875 SCE70305.1    
 GA0070614_5151   Micromonospora coxensis DSM 45161 SCG73108.1    
 GA0070609_3788   Micromonospora echinaurantiaca DSM 43904 SCG62240.1    
 GA0070610_3661   Micromonospora echinofusca DSM 43913 SCG17350.1    
 GA0070618_0026   Micromonospora echinospora DSM 43816 SCE66845.1    
 GA0070620_2251   Micromonospora krabiensis DSM 45344 SBV26757.1    
 GA0070621_0691   Micromonospora narathiwatensis DSM 45248 SBT39484.1    
 GA0074696_4466   Micromonospora purpureochromogenes DSM 43821 SCF32701.1    
 GA0070623_1716   Micromonospora rifamycinica DSM 44983 SCG49547.1    
 GA0074704_2690   Micromonospora siamensis DSM 45097 SCG51761.1    
 ML5_2300   Micromonospora sp. L5 ADU07822.1 D3C6Q5  
 CO540_08705   Micromonospora sp. WMMA2032 ATO13900.1    
 GA0070619_1501   Micromonospora zamorensis DSM 45600 SCG44077.1    
 GA0070619_1410   Micromonospora zamorensis DSM 45600 SCG43554.1    
 A7982_03337   Minicystis rosea DSM 24000 APR77990.1    
 SAMN05216490_0574   Mucilaginibacter mallensis MP1X4 SDS11895.1    
 MFUL124B02_14345   Myxococcus fulvus 124B02 AKF80604.1    
 LILAB_32135   Myxococcus fulvus HW-1 AEI68308.1    
 A176_002053   Myxococcus hansupus AKQ65141.1    
 MYMAC_004732   Myxococcus macrosporus DSM 14697 ATB49094.1    
 MYSTI_02614   Myxococcus stipitatus DSM 14675 AGC43930.1    
 MXAN_4837 (CelA)   Myxococcus xanthus DK 1622 ABF91186.1 Q1D2X5  
 CelA   Myxococcus xanthus DK101 CAA54086.1
AAW97562.1
Q50901  
 Niako_2653 (fragment)   Niastella koreensis GR20-10 AEV98992.1    
 CGQ36_26305   Nocardiopsis dassonvillei HZNU_N_1 ASU60850.1    
 A9R04_26145   Nocardiopsis dassonvillei NOCA502F APC37939.1    
 Ndas_4558   Nocardiopsis dassonvillei subsp. dassonvillei DSM 43111 ADH69945.1 D7AY67  
 DDD_2357   Nonlabens dokdonensis DSW-6 AGC77484.1    
 NMS_0139   Nonlabens marinus S1-08 BAO54148.1    
 NMS_0139   Nonlabens marinus S1-08 BAO54148.1    
 CW736_08070   Nonlabens sp. MB-3u-79 AUC79340.1    
 BN4615_P291   Nonomuraea gerenzanensis ATCC 39727 SBO90777.1    
 BN4615_P6392   Nonomuraea gerenzanensis ATCC 39727 SBO96876.1    
 BN4615_P7946   Nonomuraea gerenzanensis ATCC 39727 SBO98430.1    
 BKM31_11635   Nonomuraea sp. ATCC 55076 AQZ62037.1    
 BKM31_47795   Nonomuraea sp. ATCC 55076 AQZ68154.1    
 CW732_15140   Olleya sp. Bg11-27 AUC76937.1    
 CW732_15140   Olleya sp. Bg11-27 AUC76937.1    
 OPIT5_18400   Opitutaceae bacterium TAV5 AHF94509.1    
 OPIT5_19195   Opitutaceae bacterium TAV5 AHF92037.1    
 Oter_2143   Opitutus terrae PB90-1 ACB75426.1    
 Oter_1826   Opitutus terrae PB90-1 ACB75109.1 B1ZWZ8  
 Oter_3823   Opitutus terrae PB90-1 ACB77098.1 B1ZZ26  
 Oter_0030   Opitutus terrae PB90-1 ACB73322.1 B1ZWJ8  
 PBOR_27925   Paenibacillus borealis DSM 13188 AIQ60350.1    
 PBOR_27925   Paenibacillus borealis DSM 13188 AIQ60350.1    
 PBOR_27925   Paenibacillus borealis DSM 13188 AIQ60350.1    
 PBOR_27925   Paenibacillus borealis DSM 13188 AIQ60350.1    
 B9T62_24450   Paenibacillus donghaensis KCTC 13049 ASA23665.1    
 B9T62_24450   Paenibacillus donghaensis KCTC 13049 ASA23665.1    
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