Carbohydrate-Binding Module Family 4

Activities in FamilyModules of approx. 150 residues found in bacterial enzymes. Binding of these modules has been demonstrated with xylan, β-1,3-glucan, β-1,3-1,4-glucan, β-1,6-glucan and amorphous cellulose but not with crystalline cellulose.
3D Structure Statusβ-sandwich
NotePreviously known as cellulose-binding domain family IV (CBD IV).
External resourcesHOMSTRAD;
Statistics GenBank accession (484); Uniprot accession (125); PDB accession (18); 3D entries (10); cryst (4)
All (650) Archaea (2) Bacteria (640) unclassified (8) Structure (10 - 4 cryst) Characterized (52)
| 1 | 2 | 3 | 4 | 5 | 6 | 7 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D
 BN85307940 (LicA)   Acholeplasma brassicae O502 CCV65815.1    
 BN85307940 (LicA)   Acholeplasma brassicae O502 CCV65815.1    
 AHOG_07020 (Cel1)   Actinoalloteichus hoggarensis DSM 45943 ASO19052.1    
 TL08_07570   Actinoalloteichus hymeniacidonis HPA177(T) (=DSM 45092(T)) AOS62332.1    
 UA74_07310   Actinoalloteichus sp. ADI127-7 APU13531.1    
 UA75_07305   Actinoalloteichus sp. GBA129-24 APU19480.1    
 SAMN04489716_0458   Actinoplanes derwentensis DSM 43941 SDS29158.1    
 AFR_25825   Actinoplanes friuliensis DSM 7358 AGZ43429.1    
 AMIS_48350   Actinoplanes missouriensis 431 BAL90055.1    
 L083_3175 (Cel1)   Actinoplanes sp. N902-109 AGL16685.1    
 L083_3175 (Cel1)   Actinoplanes sp. N902-109 AGL16685.1    
 ACWT_2656   Actinoplanes sp. SE50 ATO82071.1    
 ACPL_2890 (Cel1)   Actinoplanes sp. SE50/110 AEV83785.1
SLL99479.1
   
 Amir_5339   Actinosynnema mirum DSM 43827 ACU39159.1 C6W987  
 CNX65_26585   Actinosynnema pretiosum X47 ATE56400.1    
 CDL62_16845   Alkalitalea saponilacus DSM 24412 ASB50696.1    
 CDL62_06375   Alkalitalea saponilacus DSM 24412 ASB48782.1    
 CDL62_12355   Alkalitalea saponilacus DSM 24412 ASB49870.1    
 CDL62_01845   Alkalitalea saponilacus DSM 24412 ASB47982.1    
 AXY_21140   Amphibacillus xylanus NBRC 15112 BAM48246.1    
 AXY_21140   Amphibacillus xylanus NBRC 15112 BAM48246.1    
 AXY_21140   Amphibacillus xylanus NBRC 15112 BAM48246.1    
 B737_7274   Amycolatopsis mediterranei RB AGT87935.1    
 RAM_37965   Amycolatopsis mediterranei S699 AEK46060.1
AFO80807.1
   
 AMED_7385   Amycolatopsis mediterranei U32 ADJ49099.1 D8I9D2  
 SAMN00777080_3692   Aquiflexum balticum DSM 16537 SMD45051.1    
 SAMN00777080_3692   Aquiflexum balticum DSM 16537 SMD45051.1    
 SAMN00777080_1515   Aquiflexum balticum DSM 16537 SMD42944.1    
 SAMN04489747_0990   Auraticoccus monumenti MON 2.2 SDD45976.1    
 SAMN04489747_0990   Auraticoccus monumenti MON 2.2 SDD45976.1    
 SAMN04489747_0990   Auraticoccus monumenti MON 2.2 SDD45976.1    
 BBEV_0371 (Glcα-2)   Bacillus beveridgei MLTeJB AOM81765.1    
 BBEV_0371 (Glcα-2)   Bacillus beveridgei MLTeJB AOM81765.1    
 BBEV_0314 (Glcα-1)   Bacillus beveridgei MLTeJB AOM81708.1    
 BBEV_0314 (Glcα-1)   Bacillus beveridgei MLTeJB AOM81708.1    
 Bcell_0524   Bacillus cellulosilyticus DSM 2522 ADU28806.1 D2LW92  
 Bcell_0524   Bacillus cellulosilyticus DSM 2522 ADU28806.1 D2LW92  
 Bcell_0524   Bacillus cellulosilyticus DSM 2522 ADU28806.1 D2LW92  
 Bcell_0683   Bacillus cellulosilyticus DSM 2522 ADU28965.1 D2M1H7  
 Bcell_0683   Bacillus cellulosilyticus DSM 2522 ADU28965.1 D2M1H7  
 Bcell_0683   Bacillus cellulosilyticus DSM 2522 ADU28965.1 D2M1H7  
 Bcell_0683   Bacillus cellulosilyticus DSM 2522 ADU28965.1 D2M1H7  
 Bcell_0683   Bacillus cellulosilyticus DSM 2522 ADU28965.1 D2M1H7  
 Bcell_0690   Bacillus cellulosilyticus DSM 2522 ADU28972.1    
 Bcell_0690   Bacillus cellulosilyticus DSM 2522 ADU28972.1    
 Bcell_0690   Bacillus cellulosilyticus DSM 2522 ADU28972.1    
 Bcell_1280   Bacillus cellulosilyticus DSM 2522 ADU29545.1 D2LZD8  
 B4U37_18140   Bacillus horikoshii 20a ART77832.1    
 B4U37_18140   Bacillus horikoshii 20a ART77832.1    
 B4U37_18140   Bacillus horikoshii 20a ART77832.1    
 B4U37_18140   Bacillus horikoshii 20a ART77832.1    
 BleG1_3567   Bacillus lehensis G1 AIC96114.1    
 BleG1_3567   Bacillus lehensis G1 AIC96114.1    
 BpOF4_17975   Bacillus pseudofirmus OF4 ADC51636.1 D3FS01  
 BpOF4_17975   Bacillus pseudofirmus OF4 ADC51636.1 D3FS01  
 BpOF4_17975   Bacillus pseudofirmus OF4 ADC51636.1 D3FS01  
 BpOF4_17975   Bacillus pseudofirmus OF4 ADC51636.1 D3FS01  
 A9C19_15435   Bacillus weihaiensis Alg07 APH06018.1    
 A9C19_15435   Bacillus weihaiensis Alg07 APH06018.1    
 A9C19_15435   Bacillus weihaiensis Alg07 APH06018.1    
 A9C19_15435   Bacillus weihaiensis Alg07 APH06018.1    
 endo-β-1,4-xylanase (BiXyn10C;BACINT_04197) (Xyn10C) 3.2.1.8 Bacteroides intestinalis DSM 17393 EDV05054.1    
 Bovatus_01728 (XynA)   Bacteroides ovatus ATCC 8483 ALJ46369.1    
 Bovatus_01728 (XynA)   Bacteroides ovatus ATCC 8483 ALJ46369.1    
 BACOV975_04518   Bacteroides ovatus V975 SCV10724.1    
 BACOV975_04518   Bacteroides ovatus V975 SCV10724.1    
 BXY_29300   Bacteroides xylanisolvens XB1A CBK67953.1 D6D0K1  
 Belba_2164   Belliella baltica DSM 15883 AFL84731.1    
 ADH70_018975   Blautia coccoides YL58 ASU30685.1    
 ADH70_018975   Blautia coccoides YL58 ASU30685.1    
 ADH70_018975   Blautia coccoides YL58 ASU30685.1    
 A4V09_20360   Blautia sp. YL58 ANU77877.1    
 A4V09_20360   Blautia sp. YL58 ANU77877.1    
 A4V09_20360   Blautia sp. YL58 ANU77877.1    
 CIY_16480   Butyrivibrio fibrisolvens 16/4 CBK74410.1 D4J2S1  
 CIY_07040   Butyrivibrio fibrisolvens 16/4 CBK73606.1 D4J0G7  
 CIY_07040   Butyrivibrio fibrisolvens 16/4 CBK73606.1 D4J0G7  
 Calhy_0060   Caldicellulosiruptor hydrothermalis 108 ADQ05822.1 E4Q9F1  
 Calhy_0060   Caldicellulosiruptor hydrothermalis 108 ADQ05822.1 E4Q9F1  
 Calhy_0060   Caldicellulosiruptor hydrothermalis 108 ADQ05822.1 E4Q9F1  
 Calhy_0060   Caldicellulosiruptor hydrothermalis 108 ADQ05822.1 E4Q9F1  
 Calhy_0060   Caldicellulosiruptor hydrothermalis 108 ADQ05822.1 E4Q9F1  
 Calkro_0072   Caldicellulosiruptor kronotskyensis 2002 ADQ44988.1 E4SCD3  
 Calkro_0072   Caldicellulosiruptor kronotskyensis 2002 ADQ44988.1 E4SCD3  
 Calkro_0072   Caldicellulosiruptor kronotskyensis 2002 ADQ44988.1 E4SCD3  
 Calkro_0072   Caldicellulosiruptor kronotskyensis 2002 ADQ44988.1 E4SCD3  
 Calkro_0072   Caldicellulosiruptor kronotskyensis 2002 ADQ44988.1 E4SCD3  
 COB47_0076   Caldicellulosiruptor obsidiansis OB47 ADL41452.1 D9TGW6  
 COB47_0076   Caldicellulosiruptor obsidiansis OB47 ADL41452.1 D9TGW6  
 COB47_0076   Caldicellulosiruptor obsidiansis OB47 ADL41452.1 D9TGW6  
 COB47_0076   Caldicellulosiruptor obsidiansis OB47 ADL41452.1 D9TGW6  
 COB47_0076   Caldicellulosiruptor obsidiansis OB47 ADL41452.1 D9TGW6  
 Csac_2548 (fragment)   Caldicellulosiruptor saccharolyticus DSM 8903 ABP68121.1
ABP68121.2
A4XMI6  
 Csac_2548 (fragment)   Caldicellulosiruptor saccharolyticus DSM 8903 ABP68121.1
ABP68121.2
A4XMI6  
 Csac_2548 (fragment)   Caldicellulosiruptor saccharolyticus DSM 8903 ABP68121.1
ABP68121.2
A4XMI6  
 Csac_2548 (fragment)   Caldicellulosiruptor saccharolyticus DSM 8903 ABP68121.1
ABP68121.2
A4XMI6  
 Csac_2548 (fragment)   Caldicellulosiruptor saccharolyticus DSM 8903 ABP68121.1
ABP68121.2
A4XMI6  
 endo-1,3(4)-β-glucanase, partial (Lam16A)   Caldicellulosiruptor sp. F32 ALP73406.1    
 endo-1,3(4)-β-glucanase, partial (Lam16A)   Caldicellulosiruptor sp. F32 ALP73406.1    
 endo-1,3(4)-β-glucanase, partial (Lam16A)   Caldicellulosiruptor sp. F32 ALP73406.1    
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