Carbohydrate-Binding Module Family 22

Activities in FamilyA xylan binding function has been demonstrated in several cases and affinity with mixed β-1,3/β-1,4-glucans in one. In several cases a thermostabilizing effect has also been seen.
3D Structure Statusβ-sandwich
NoteFormerly known as X6 modules or thermostabilizing domains
External resourcesHOMSTRAD;
Commercial Enzyme Provider(s)NZYTech; PROZOMIX;
Statistics GenBank accession (712); Uniprot accession (189); PDB accession (13); 3D entries (4); cryst (0)
| 1 | 2 |
Bacteria
Protein Name EC#OrganismGenBank UniprotPDB/3D
 xylanase E (XynE) 3.2.1.8 Aeromonas caviae ME-1 BAA34091.1 Q9Z485  
 β-1,4-xylanase (ORF2504) (Xyn10A) 3.2.1.- Caldanaerobius polysaccharolyticus KMCJ AFQ62797.1    
 β-1,4-xylanase (ORF2504) (Xyn10A) 3.2.1.- Caldanaerobius polysaccharolyticus KMCJ AFQ62797.1    
 xylanase (XynA) 3.2.1.8 Caldibacillus cellulovorans AAF61649.1 Q9L8L8  
 xylanase A (XynA;Athe_0618) 3.2.1.8 Caldicellulosiruptor bescii DSM 6725 CAA93627.1
ACM59742.1
Q59150  
 xylanase A (XynA;Athe_0618) 3.2.1.8 Caldicellulosiruptor bescii DSM 6725 CAA93627.1
ACM59742.1
Q59150  
 xylanase (Cb193;CbXyn10A;Athe_0183) 3.2.1.8 Caldicellulosiruptor bescii DSM 6725 ACM59335.1 B9MMA3  
 xylanase (Cb193;CbXyn10A;Athe_0183) 3.2.1.8 Caldicellulosiruptor bescii DSM 6725 ACM59335.1 B9MMA3  
 exo-xylanase (Calkro_0402) 3.2.1.- Caldicellulosiruptor kronotskyensis 2002 ADQ45302.1 E4SE15  
 exo-xylanase (Calkro_0402) 3.2.1.- Caldicellulosiruptor kronotskyensis 2002 ADQ45302.1 E4SE15  
 exo-xylanase (Calkro_0402) 3.2.1.- Caldicellulosiruptor kronotskyensis 2002 ADQ45302.1 E4SE15  
 β-xylosidase / α-L-arabinofuranosidase (XynF) 3.2.1.37
3.2.1.55
Caldicellulosiruptor saccharolyticus AAB87371.1 O30426  
 xylanase E (XynE) 3.2.1.8 Caldicellulosiruptor saccharolyticus AAB87372.1 O30427  
 xylanase E (XynE) 3.2.1.8 Caldicellulosiruptor saccharolyticus AAB87372.1 O30427  
 XynI 3.2.1.8 Caldicellulosiruptor saccharolyticus AAB87379.1 O30421  
 XynI 3.2.1.8 Caldicellulosiruptor saccharolyticus AAB87379.1 O30421  
 xylanase (XynE;Csac_2410) 3.2.1.8 Caldicellulosiruptor saccharolyticus DSM 8903 ABP67987.1 A4XM52  
 xylanase (XynE;Csac_2410) 3.2.1.8 Caldicellulosiruptor saccharolyticus DSM 8903 ABP67987.1 A4XM52  
 bifunctional α-L-arabinofuranosidase (XynF;Csac_2411) 3.2.1.55 Caldicellulosiruptor saccharolyticus DSM 8903 ABP67988.1 A4XM53  
 xylanase (XynB;Ca_GH10) 3.2.1.8 Caldicellulosiruptor sp. F32 AFO70073.1    
 xylanase (XynB;Ca_GH10) 3.2.1.8 Caldicellulosiruptor sp. F32 AFO70073.1    
 xylanase B (XynB) 3.2.1.8 Caldicellulosiruptor sp. Rt69B.1 AAB95325.1 O52373  
 xylanase B (XynB) 3.2.1.8 Caldicellulosiruptor sp. Rt69B.1 AAB95325.1 O52373  
 xylanase B (XynB) 3.2.1.8 Caldicellulosiruptor sp. Rt69B.1 AAB95325.1
CAA03086.1
CAE84530.1
O52373  
 xylanase C (XynC) (multidomain protein) 3.2.1.8 Caldicellulosiruptor sp. Rt69B.1 AAB95326.1
CAA03087.1
CAE84531.1
O52374  
 xylanase C (XynC) (multidomain protein) 3.2.1.8 Caldicellulosiruptor sp. Rt69B.1 AAB95326.1 O52374  
 xylanase A (XynA) 3.2.1.8 Caldicellulosiruptor sp. Rt8B.4 AAB42044.1 P40944  
 xylanase A (XynA) 3.2.1.8 Caldicellulosiruptor sp. Rt8B.4 AAB42044.1 P40944  
 bifunctional β-1,4-xylanase / α-L-arabinosidase (XynA) 3.2.1.55
3.2.1.8
Caldicellulosiruptor sp. Tok7B.1 AAD30363.1 Q9X3P5  
 bifunctional β-1,4-xylanase / α-L-arabinosidase (XynA) 3.2.1.55
3.2.1.8
Caldicellulosiruptor sp. Tok7B.1 AAD30363.1 Q9X3P5  
 xylanase C (XynC;XylC) 3.2.1.8 Cellulomonas fimi ATCC 484 CAA90745.1 Q59278  
 xylanase B (XynB) 3.2.1.8 Clostridium cellulovorans ATCC 35296 AAT37531.1 Q6J286  
 xylanase A (XynA) (Xyn10A) 3.2.1.8 Clostridium josui FERM P-9684 BAB20931.1 Q9F1V3  
 xylanase A (XynA) (Xyn10A) 3.2.1.8 Clostridium josui FERM P-9684 BAB20931.1 Q9F1V3  
 xylanase C / β-1,3-1,4-glucanase (XynC) (Xyn10B) 3.2.1.8 Clostridium stercorarium BAA82143.1 Q9XDV5  
 xylanase C / β-1,3-1,4-glucanase (XynC) (Xyn10B) 3.2.1.8 Clostridium stercorarium BAA82143.1 Q9XDV5  
 celloxylanase (CelX;XynC) (Xyn10C) 3.2.1.8 Clostridium stercorarium NCIMB 11745 CAD48314.1 Q8GJ37  
 celloxylanase (CelX;XynC) (Xyn10C) 3.2.1.8 Clostridium stercorarium NCIMB 11745 CAD48314.1 Q8GJ37  
 xylanase (Dtur_1715) 3.2.1.- Dictyoglomus turgidum DSM 6724 ACK42987.1 B8E3B3  
 xylanase (Dtur_1715) 3.2.1.- Dictyoglomus turgidum DSM 6724 ACK42987.1 B8E3B3  
 xylanase C / 10 C (XynC;X-K) (Xyn10C) 3.2.1.8 Paenibacillus barcinonensis BP-23 CAA07173.1 O69230 4XUN[A,B,C]
4XUP[A,B,D,F]
4XUQ[A,B,C]
4XUR[A,B,C]
4XUT[A,B,C]
 xylanase C / 10 C (XynC;X-K) (Xyn10C) 3.2.1.8 Paenibacillus barcinonensis BP-23 CAA07173.1 O69230 4XUO[A,B]
4XUP[A,B,C,D,E,F]
 xylanase 10A (Xyn10A) 3.2.1.- Paenibacillus curdlanolyticus B-6 ABZ80916.1 B1A3N2  
 xylanase 10A (Xyn10A) 3.2.1.- Paenibacillus curdlanolyticus B-6 ABZ80916.1 B1A3N2  
 xylanase 10A (Xyn10A) 3.2.1.- Paenibacillus curdlanolyticus B-6 ABZ80916.1 B1A3N2  
 endo-xylanase (XynC) 3.2.1.8 Paenibacillus sp. DG-22 AIE12545.1    
 endo-xylanase (XynC) 3.2.1.8 Paenibacillus sp. DG-22 AIE12545.1    
 methylglucuronoxylan xylanase A / xylanase A1 (XynA1;PbXynA1;XynA;Pjdr2_0221) 3.2.1.8 Paenibacillus sp. JDR-2 CAI79477.1
ACS98901.1
C6CRV0
Q53I45
 
 methylglucuronoxylan xylanase A / xylanase A1 (XynA1;PbXynA1;XynA;Pjdr2_0221) 3.2.1.8 Paenibacillus sp. JDR-2 CAI79477.1
ACS98901.1
C6CRV0
Q53I45
 
 methylglucuronoxylan xylanase A / xylanase A1 (XynA1;PbXynA1;XynA;Pjdr2_0221) 3.2.1.8 Paenibacillus sp. JDR-2 CAI79477.1
ACS98901.1
C6CRV0
Q53I45
 
 xylanase 5 (Xyn5) 3.2.1.8 Paenibacillus sp. W-61 BAC45001.1 Q8GHJ4  
 xylanase 5 (Xyn5) 3.2.1.8 Paenibacillus sp. W-61 BAC45001.1 Q8GHJ4  
 xylanase (KRICT PX3;KRICT-PX3;Xy1;HPL003_05890;Pae1223) 3.2.1.8 Paenibacillus terrae HPL-003 AET57943.1
ADX97440.1
   
 xylanase (KRICT PX3;KRICT-PX3;Xy1;HPL003_05890;Pae1223) 3.2.1.8 Paenibacillus terrae HPL-003 AET57943.1
ADX97440.1
   
 xylanase (Theth_1635) (Xyn10A) 3.2.1.8 Pseudothermotoga thermarum DSM 5069 AEH51685.1    
 xylanase (Theth_1635) (Xyn10A) 3.2.1.8 Pseudothermotoga thermarum DSM 5069 AEH51685.1    
 xylanase (Theth_1635) (Xyn10A) 3.2.1.8 Pseudothermotoga thermarum DSM 5069 AEH51685.1    
 xylanase (XylL) 3.2.1.8 Psychrobacter sp. 2-17 AFA53084.1    
 xylanase Y (XynY;Cthe_0912) (Xyn10B) 3.2.1.8 Ruminiclostridium thermocellum ATCC 27405 ABN52146.1 A3DDW7  
 xylanase Y (XynY;Cthe_0912) (Xyn10B) 3.2.1.8 Ruminiclostridium thermocellum ATCC 27405 ABN52146.1 A3DDW7  
 xylanase C (XynC;Cthe_1838) (Xyn10C) 3.2.1.8 Ruminiclostridium thermocellum ATCC 27405 ABN53059.1 A3DGI0  
 β-1,4-xylanase (XynD;Cthe_2590) (Xyn10D) 3.2.1.8 Ruminiclostridium thermocellum ATCC 27405 ABN53790.1 A3DIL1  
 xylanase C (XynC) (Xyn10C) 3.2.1.8 Ruminiclostridium thermocellum F1 BAA21516.2 O32374  
 β-1,4-xylanase (XynD) (Xyn10D) 3.2.1.8 Ruminiclostridium thermocellum F7 CAE51307.1 Q70DK4  
 xylanase Y / feruloyl esterase (XynY;LX5) (Xyn10B) 3.1.1.73
3.2.1.8
Ruminiclostridium thermocellum YS CAA58242.1
AAR39814.1
P51584 1DYO[A,B]
1H6X[A]
1H6Y[A,B]
 xylanase Y / feruloyl esterase (XynY;LX5) (Xyn10B) 3.1.1.73
3.2.1.8
Ruminiclostridium thermocellum YS CAA58242.1
AAR39814.1
P51584 2W5F[A,B]
2WYS[A,B]
2WZE[A,B]
 xylanase A (XynA) 3.2.1.8 Ruminococcus albus AAA85198.1 Q52644  
 xylanase B (XynB) 3.2.1.8 Ruminococcus albus BAB39493.1 Q9AJB4  
 endo-β-1,4-xylanase 11D (Xyn11D) 3.1.1.72
3.2.1.8
Ruminococcus albus 8 AEE64769.1
ZP_08160424.1
   
 endo-β-1,4-xylanase 11E (Xyn11E) 3.2.1.8 Ruminococcus albus 8 AEE64770.1
ZP_08158180.1
   
 endo-β-1,4-xylanase 10B (Xyn10B) 3.2.1.8 Ruminococcus albus 8 AEE64768.1
ZP_08159615.1
   
 endo-β-1,4-xylanase 10B (Xyn10B) 3.2.1.8 Ruminococcus albus 8 AEE64768.1
ZP_08159615.1
   
 endo-β-1,4-xylanase 11F (Xyn11F) 3.2.1.8 Ruminococcus albus 8 AEE64771.1
ZP_08157788.1
   
 xylanase (GH10A;RUM_03400) (Xyn10A) 3.2.1.8 Ruminococcus champanellensis 18P13 = JCM 17042 CBL16579.1 D4LAD4  
 bifunctional xylanase α-L-arabinofuranosidase (GH10B;GH43E;RUM_15940) (Xyn10B-Abf43E) 3.2.1.55
3.2.1.8
Ruminococcus champanellensis 18P13 = JCM 17042 CBL17682.1 D4LDI7  
 bifunctional xylanase α-L-arabinofuranosidase (GH10B;GH43E;RUM_15940) (Xyn10B-Abf43E) 3.2.1.55
3.2.1.8
Ruminococcus champanellensis 18P13 = JCM 17042 CBL17682.1 D4LDI7  
 xylanase (GH30A-CE;RUM_18530) (Xyn30A) 3.2.1.8 Ruminococcus champanellensis 18P13 = JCM 17042 CBL17903.1 D4LE58  
 xylanase (GH11A-CE;RUM_23130) (Xyn11A) 3.2.1.8 Ruminococcus champanellensis 18P13 = JCM 17042 CBL18305.1 D4LFB0  
 xylanase (GH11A-CE;RUM_23130) (Xyn11A) 3.2.1.8 Ruminococcus champanellensis 18P13 = JCM 17042 CBL18305.1 D4LFB0  
 xylanase B (XynB) 3.1.1.72
3.2.1.8
Ruminococcus flavefaciens 17 CAA84537.1 Q52753  
 xylanase D / lichenase (XynD) 3.2.1.73
3.2.1.8
Ruminococcus flavefaciens 17 AAB26620.1
CAB51934.1
Q53317
Q9S310
 
 endoxylanase E (XynE) 3.2.1.8 Ruminococcus flavefaciens 17 CAB93667.1 Q9L3K3  
 xylanase 1 (Xyn1) 3.2.1.8 Ruminococcus sp. CAA90271.1
AAR39816.1
Q59790  
 xylanase (XynA) 3.2.1.8 Thermoanaerobacterium aotearoense SCUT27 AKP07634.1    
 xylanase (XynA) 3.2.1.8 Thermoanaerobacterium aotearoense SCUT27 AKP07634.1    
 xylanase A (XynA) 3.2.1.8 Thermoanaerobacterium saccharolyticum B6A-RI AAA21812.1 P36917  
 xylanase A (XynA) 3.2.1.8 Thermoanaerobacterium saccharolyticum B6A-RI AAA21812.1 P36917  
 endo-β-1,4-xylanase (XynA;XylA;Tsac_1459) 3.2.1.8 Thermoanaerobacterium saccharolyticum JW/SL-YS485 AFK86466.1
AAC43719.1
Q60043  
 endo-β-1,4-xylanase (XynA;XylA;Tsac_1459) 3.2.1.8 Thermoanaerobacterium saccharolyticum JW/SL-YS485 AFK86466.1
AAC43719.1
Q60043  
 xylanase X (XynX) 3.2.1.8 Thermoanaerobacterium sp. AAA23227.1 P38535  
 xylanase (XynA) (salt-tolerant) 3.2.1.8 Thermoanaerobacterium sp. NTOU2 ADB23440.1    
 xylanase (XynA) (salt-tolerant) 3.2.1.8 Thermoanaerobacterium sp. NTOU2 ADB23440.1    
 xylanase (Xyn;XYN;Tthe_0992) 3.2.1.8 Thermoanaerobacterium thermosaccharolyticum DSM 571 ADL68522.1
YP_003851606.1
D9TMZ9  
 xylanase A (XynA) 3.2.1.8 Thermoanaerobacterium thermosulfurigenes EM1 AAB08046.1 Q60046  
 xylanase A (XynA) 3.2.1.8 Thermoanaerobacterium thermosulfurigenes EM1 AAB08046.1 Q60046  
 xylanase A (XynA;Tm0061) (Xyl10A) 3.2.1.8 Thermotoga maritima MSB8 AAD35155.1
AGL48984.1
AHD18169.1
AKE27869.1
CAA86406.1
NP_227877.1
G4FGX6
Q60037
 
 xylanase A (XynA;Tm0061) (Xyl10A) 3.2.1.8 Thermotoga maritima MSB8 AAD35155.1
AGL48984.1
AHD18169.1
AKE27869.1
CAA86406.1
NP_227877.1
G4FGX6
Q60037
 
 xylanase A (XynA;CTN_0632) 3.2.1.8 Thermotoga neapolitana DSM 4359 ACM22808.1
CAA87069.1
B9K775
Q60042
 
 xylanase A (XynA;CTN_0632) 3.2.1.8 Thermotoga neapolitana DSM 4359 ACM22808.1
CAA87069.1
B9K775
Q60042
 
 xylanase B (XynB) 3.2.1.8 Thermotoga sp. strain FjSS3-B.1 AAD32593.1
AAD32593.2
Q9R6T4  
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