Carbohydrate-Binding Module Family 13

Known ActivitiesModules of approx. 150 residues which always appear as a threefold internal repeat. The only apparent exception to this, xylanase II of Actinomadura sp. FC7 (GenBank U08894), is in fact not completely sequenced. These modules were first identified in several plant lectins such as ricin or agglutinin of Ricinus communis which bind galactose residues. The three-dimensional structure of a plant lectin has been determined and displays a pseudo-threefold symmetry in accord with the observed sequence threefold repeat. These modules have since been found in a number of other proteins of various functions including glycoside hydrolases and glycosyltransferases. While in the plant lectins this module binds mannose, binding to xylan has been demonstrated in the Streptomyces lividans xylanase A and arabinofuranosidase B. Binding to GalNAc has been shown for the corresponding module of GalNAc transferase 4. For the other proteins, the binding specificity of these modules has not been established. The pseudo three-fold symmetry of the CBM13 module has now been confirmed in the 3-D structure of the intact, two-domain, xylanase of Streptomyces olivaceoviridis.
3D Structure Statusb-trefoil
NotePreviously known as cellulose-binding domain family XIII (CBD XIII).
External resourcesHOMSTRAD;
Statistics GenBank accession (2377); Uniprot accession (891); PDB accession (85); 3D entries (41); cryst (1)
All (1793) Archaea (13) Bacteria (1361) Eukaryota (401) Viruses (10) unclassified (8) Structure (41 - 1 cryst) Characterized (87)
Bacteria
Protein Name EC#OrganismGenBankUniprotPDB/3D
 xylanase II (XylII) (fragment) 3.2.1.8 Actinomadura sp. FC7 AAA17888.1
AAO97631.1
AAQ80272.1
Q59139  
 β-agarase (AgaB34) 3.2.1.81 Agarivorans albus YKW-34 ABW77762.1 A8W969  
 β-agarase (AgaJA3;Hz4) 3.2.1.81 Agarivorans sp. JA-1 / HZ105 ADY17921.1    
 β-agarase (AgaA) 3.2.1.81 Agarivorans sp. LQ48 ACM50513.1 C9WIW7  
 β-1,3-glucanase (GlcI) 3.2.1.39 Arthrobacter sp. YCWD3 BAA04892.1 Q59146  
 β-1,3(4)-glucanase M (BglM) 3.2.1.39
3.2.1.6
3.2.1.73
Bacillus circulans IAM1165 BAC06195.1 Q8KKH3  
 pectate lyase 4.2.2.2 Bacillus sp. AAI12 AAE59747.1
AAS29294.1
   
 feruloyl esterase I (FaeI;CGSCsYakCAS_18248) (Fae1A) 3.1.1.73 Cellulosilyticum ruminicola CGMCC 1.5065 ACZ98598.1 D2KFJ6  
 β-1,3-glucanase (BglI) 3.2.1.39 Cellulosimicrobium cellulans 73 14 / ATCC21606 / DSM 10297 AAA25520.1 P22222  
 β-1,3-glucanase II (BglII) 3.2.1.39 Cellulosimicrobium cellulans LL-G109 / DSM10297 / DK-1 AAC38290.1
AEK80404.1
O68641  
 exo-β-1,3-galactanase (Ct1,3Gal43A;Cthe_0661) (1,3Gal43A) 3.2.1.145 Clostridium thermocellum ATCC 27405 ABN51896.1 A3DD67 3VSF[A,B,C,D,E,F]
3VSZ[A,B,C,D,E,F]
3VT0[A,B,C,D,E,F]
3VT1[A,B,C,D,E,F]
3VT2[A,B,C,D,E,F]
 xylanase (M6;CtM6;Cthe_2193) (arabinoxylan-specific) (Xyl5A) 3.2.1.- Clostridium thermocellum ATCC 27405 ABN53395.1 A3DHG6  
 β-agarase (AgaY) (extracellular) 3.2.1.81 Janthinobacterium sp. SY12 ABO93616.2 A4UMV9  
 pectate lyase B (PelB) 4.2.2.2 Pseudoalteromonas haloplanktis 505 AAF86343.1
AAF86343.2
Q9KGY6  
 β-agarase (AgaA) (extracellular) 3.2.1.81 Pseudoalteromonas sp. CY24 AAN39119.2 Q8GB62  
 β-agarase (AgaA6) 3.2.1.81 Pseudoalteromonas sp. JT-6 ACF40223.1 C0J1J2  
 β-agarase (PagA) 3.2.1.81 Pseudomonas sp. BK1 SK38 / BK1      
 β-porphyranase (AagA) 3.2.1.178 Pseudomonas sp. ND137 BAB79291.1 Q8VUT1  
 alginate lyase / poly(β-mannuronate) lyase (Sde_2547) (Alg7D) 4.2.2.3 Saccharophagus degradans 2-40 ABD81807.1
ACK10570.1
ACK10582.1
Q21HM2  
 endo-β-1,3-xylanase (Xyl30) 3.2.1.32 Streptomyces avermitilis CECT3339 AAD32560.1 Q9X584  
 exo-β-1,3-galactanase (Sa1,3Gal43A;SAV2109) (1,3Gal43A) 3.2.1.145 Streptomyces avermitilis MA-4680 BAC69820.1
NP_823285.1
   
 β-L-arabinopyranosidase (AgaA2;SaArap27A;SAV_2186) (Arap27A) 3.2.1.88 Streptomyces avermitilis MA-4680 BAC69897.1
NP_823362.1
  3A21[A,B]
3A22[A,B]
3A23[A,B]
 bifunctional xylanase / α-L-arabinofuranosidase (Xyl23) 3.2.1.55
3.2.1.8
Streptomyces chattanoogensis UAH23 AAD32559.2
ABC05324.1
Q9X583  
 xylanase A (XlnA;SCO5931;SC10A5.36c) 3.2.1.8 Streptomyces coelicolor A3(2) CAA16468.1
CAD55241.1
NP_733679.1
Q8CJQ1  
 xylanase (Xyn10) 3.2.1.8 Streptomyces fradiae var. k11 ABR14270.1 A7TVD4  
 α-L-arabinofuranosidase B (AbfB) 3.2.1.55 Streptomyces lividans 1326 AAC26524.1 P96463  
 xylanase A (XlnA) (Xyn10A) 3.2.1.8 Streptomyces lividans 1326 / IAF18 AAC26525.1
AAO97633.1
P26514 1KNL[A]
1KNM[A]
1KNN[A]
1MC9[A]
 β-1,4-xylanase (XynA;FXYN;SoXyn10A) (Xyn10A) 3.2.1.8 Streptomyces olivaceoviridis E-86   Q7SI98 1ISV[A,B]
1ISW[A,B]
1ISX[A,B]
1ISY[A,B]
1ISZ[A,B]
1IT0[A,B]
1V6U[A,B]
1V6V[A,B]
1V6W[A,B]
1V6X[A,B]
2D1Z[A,B]
2D20[A,B]
2D22[A,B]
2D23[A,B]
2D24[A,B]
1XYF[A]
 exo-β-1,3-galactanase (SGalase2) 3.2.1.145 Streptomyces sp. 19(2012) AFH55135.1    
 exo-β-1,3-galactanase (SGalase1) (partial) 3.2.1.145 Streptomyces sp. 19(2012) AFH55134.1    
 endo-β-1,3-glucanase (BglS27) 3.2.1.39 Streptomyces sp. S27 ACO94508.1 C3VPJ1  
 xylanase (XynAS27) 3.2.1.8 Streptomyces sp. S27 ACF57946.1 C3RYK8  
 endo-β-1,4-xylanase (XynSW2;XynSW2A;XynSW2B) 3.2.1.8 Streptomyces sp. SWU10 BAK19338.1    
 xylanase A 3.2.1.8 Streptomyces thermocyaneoviolaceus AAF04600.1 Q9RMM5  
 xylanase I (StXI) 3.2.1.8 Streptomyces thermoviolaceus OPC-520 BAA19777.1
BAD02382.1
Q76BV3  
 chitinase (Chi35) 3.2.1.14 Streptomyces thermoviolaceus OPC-520 BAA88833.1 Q9RHU5  
 endoxylanase (XynB) (Xyn10A) 3.2.1.8 Thermopolyspora flexuosa DSM43186 / ATCC35864 AAO97629.1
AAO97630.1
AAO97632.1
AAQ80270.1
AAQ80271.1
AAQ80273.1
AAS23966.1
AAS23967.1
AAS23969.1
CAD48748.1
CAD48748.2
Q8GMV6  
 β-agarase D (AgaD) 3.2.1.81 Vibrio sp. PO-303 BAF34350.1 Q05KJ8  
 β-agarase (AgaV) 3.2.1.81 Vibrio sp. V134 ABL06969.1 A1E2B6  
Eukaryota
Protein Name EC#OrganismGenBankUniprotPDB/3D
 [Blood group A] α-N-acetylgalactosaminidase (NagA) 3.2.1.49 Acremonium sp. No.413 BAB08149.1 Q9HFB1  
 α-N-acetylgalactosaminidase / α-galactosidase (AglA;An06g00170) 3.2.1.22
3.2.1.49
Aspergillus niger CBS 513.88 CAK44933.1 A2QL72  
 α-galactosidase A (Ag1A;AglA) 3.2.1.22 Aspergillus niger N402 AAE69559.1
CAA44950.1
P28351  
 UDP-GalNAc: polypeptide N-acetylgalactosaminyltransferase 1 (GalNT1;ppGalNAcT1;GalNAc-T1) 2.4.1.41 Bos taurus AAA30532.1
AAA68489.1
AAE24810.1
AAE24816.1
NP_803485.1
Q07537  
 ZK688.8 (gly3) 2.4.1.41 Caenorhabditis elegans Bristol N2 AAA28224.3
AAC13669.1
CCD62541.1
NP_498722.1
P34678  
 Y116F11B.12 (Gly-4) 2.4.1.41 Caenorhabditis elegans Bristol N2 AAC13670.1
CAC14394.1
CBA11615.1
NP_507850.1
Q8I136
Q8I136
 
 H38K22.5 (Gly-6) 2.4.1.41 Caenorhabditis elegans Bristol N2 AAC13674.1
AAC13675.1
AAC13676.1
CAA19707.1
CAC42317.1
CAC42318.1
NP_001022644.1
O61394  
 Y46H3A.6 (gly-7) 2.4.1.41 Caenorhabditis elegans Bristol N2 AAC13677.1
AAF60620.1
CCD70652.1
NP_503512.1
O61397  
 GLY9 (Y47D3A.23) 2.4.1.41 Caenorhabditis elegans Bristol N2 AAC13679.1
CAB57897.1
CAI46621.1
NP_499451.1
Q9U2C4
Q9U2C4
 
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase T3 2.4.1.41 Cryptosporidium parvum AAQ95175.1
AAQ95175.2
Q6TBQ9  
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase T1 2.4.1.41 Cryptosporidium parvum AAQ95174.1
AAQ95174.2
Q6TBR0  
 polypeptide N-acetylgalactosaminyltransferase T4 (CG31956;AT25481p1) 2.4.1.41 Drosophila melanogaster AAN10370.1
AAN10370.2
AAN75751.1
AAQ56701.1
ACL82986.1
AER27632.1
NP_722910.1
B7Z006
Q8IA42
Q8IA42
 
 polypeptide N-acetylgalactosaminyltransferase T3 (CG4445) 2.4.1.41 Drosophila melanogaster AAD38630.1
AAF59298.1
NP_610256.1
Q9Y117  
  polypeptide N-acetylgalactosaminyltransferase T35A / l(2)35Aa (Pgant35A;CG7480) 2.4.1.41 Drosophila melanogaster AAF44871.1
AAF53391.1
AAK66862.1
AAM62405.1
AAM62406.1
AEG66944.1
NP_652069.2
F5XVG5
Q8MVS5
Q8MVS5
 
 polypeptide N-acetylgalactosaminyltransferase T7 (CG6394) 2.4.1.41 Drosophila melanogaster AAF48851.1
AAM62412.1
AAN09470.1
AAQ56704.1
AAV37008.1
NP_573301.2
NP_728178.1
Q8MV48
Q8MV48
 
 polypeptide N-acetylgalactosaminyltransferase T6 (CG2103) 2.4.1.41 Drosophila melanogaster AAF47689.1
AAF47690.1
AAL29177.1
AAQ56703.1
AGB94037.1
NP_647749.2
NP_728779.1
Q6WV16  
 polypeptide N-acetylgalactosaminyltransferase T8 (CG7297) 2.4.1.41 Drosophila melanogaster AAF49587.1
AAL48588.1
NP_648800.1
Q9VUT6  
 polypeptide N-acetylgalactosaminyltransferase T2 (CG3254) 2.4.1.41 Drosophila melanogaster AAF51113.2
AAF51113.3
AAQ22499.1
AAQ56700.1
AAS64620.1
AAS64620.2
NP_608773.1
Q6WV19  
  polypeptide N-acetylgalactosaminyltransferase T5 (CG31651) 2.4.1.41 Drosophila melanogaster AAF52218.2
AAL25377.1
AAQ56702.1
ABI31292.1
NP_608906.2
Q6WV17  
 polypeptide N-acetylgalactosaminyltransferase T1 (CG8182) 2.4.1.41 Drosophila melanogaster AAF58130.1
AAG13184.1
AAM29416.1
AAM70974.1
AAQ56699.1
NP_611043.1
NP_725472.1
Q6WV20  
 UDP-GalNAc: polypeptide N-acetylgalactosaminyltransferase 2 (GALNTL2;GALNT7) 2.4.1.41 Homo sapiens AAE72285.1
AAE73125.1
AAH14789.1
AAH30625.1
AAK63127.1
AAQ88808.1
ABA31473.1
ABA36473.1
ABA38055.1
ABA68732.1
ABA69735.1
ABA70117.1
ABA70240.1
ABC04061.1
ABC16096.1
ABE23551.1
ABE26114.1
ABE26387.1
ABH64624.1
ABH64747.1
ABH64904.1
ABH66133.1
ABH70952.1
ABH71075.1
ABH72589.1
ABH75704.1
ABH99489.1
ABI02594.1
ABI07395.1
ABI11043.1
ABI12263.1
ABJ19428.1
ABJ34423.1
ABL21883.1
ABX94001.1
ABX96294.1
ABX96604.1
ABY05068.1
ABY16007.1
ABY18804.1
ACC22268.1
ACC22839.1
ACC23718.1
ACC32509.1
ACC34084.1
ACE08795.1
ACG89706.1
ACK00002.1
ACM95112.1
ACN08875.1
ACP73867.1
ACS06493.1
ACS08649.1
ACW23860.1
BAD29961.1
BAG52928.1
CAD38585.1
CAD89983.1
CAG32963.1
CAK32357.1
NP_473451.2
F1LIP6
Q8N3T1
 
 Polypeptide GalNAc transferase T13 (contains KIAA1918) 2.4.1.41 Homo sapiens AAI01032.1
AAI01034.1
AAI01035.1
ACC22253.1
BAC54545.1
BAG54747.1
CAD44533.2
CAG32964.1
NP_443149.2
Q68VI8
Q8IUC8
 
 UDP-GalNAc: polypeptide N-acetylgalactosaminyltransferase 14 (GalNT14) 2.4.1.41 Homo sapiens AAH06269.1
AAH06269.2
AAH10659.1
AAQ89118.1
AAX88899.1
ABA66763.1
ABH82928.1
ABI02314.1
ABI12599.1
ABJ30825.1
ABJ38955.1
ABX91594.1
ABX91870.1
ABX92146.1
ABX92422.1
ABX92698.1
ABX93132.1
ABX93639.1
ABX94373.1
ABX95380.1
ABX95656.1
ABX95932.1
ABY04154.1
ABY04430.1
ABY04706.1
ABY05292.1
ABY16398.1
ABY17890.1
ABY18166.1
ABY18442.1
ACC25543.1
ACC26284.1
ACC36580.1
ACC36856.1
ACE07501.1
ACE09339.1
ACE10551.1
ACJ96114.1
ACN02380.1
ACN06790.1
ACP73505.1
ACP89033.1
ACR88983.1
ACR89535.1
ACW28625.1
BAB14227.1
BAB14795.1
BAC03634.1
BAC56889.1
BAG53701.1
BAH12861.1
CAA70505.3
CAA70505.4
CAD10224.1
CAD34770.1
CAD44543.1
CAE91230.1
CAE91506.1
NP_078848.2
Q96FL9  
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase (GalNAc-T11;ppGalNAcT11;GalNT11) 2.4.1.41 Homo sapiens AAD45821.1
ACC22266.1
BAB15105.1
BAB15338.1
BAG54116.1
BAH12901.1
CAC79625.2
CAC79625.3
CAD38481.1
NP_071370.2
Q8NCW6
Q8NCW6
 
 UDP-GalNAc:polypeptide α-N-acetylgalactosaminyltransferase T4 (GalNAc-T4;ppGalNAcT4;GALNT4) 2.4.1.41 Homo sapiens AAH36390.1
AAN94203.1
AAN94205.1
BAG35722.1
BAG60038.1
CAA69875.1
NP_003765.1
Q8N4A0
Q8N4A0
 
 UDP-GalNAc:polypeptide α-N-acetylgalactosaminyltransferase T1 (GalNAc-T1;ppGalNAcT1;GALNT1) 2.4.1.41 Homo sapiens AAC50327.1
AAH38440.1
ACC22256.1
BAF85751.1
CAA59380.1
CAD44535.1
NP_065207.2
Q10472
Q10472
 
 UDP-GalNAc:polypeptide α-N-acetylgalactosaminyltransferase T2 (GalNAc-T2;ppGalNAcT2;GalNT2) 2.4.1.41 Homo sapiens AAH41120.1
ACC22257.1
ACS13744.1
BAF82737.1
BAH12448.1
BAH13288.1
CAA59381.1
CAC00585.2
CAI22902.1
CAI23447.1
NP_004472.1
Q10471 2FFU[A]
2FFV[A,B]
 UDP-GalNAc:polypeptide α-N-acetylgalactosaminyltransferase T3 (ppGalNacT3;GALNAc-T3;GALNT3) 2.4.1.41 Homo sapiens AAI13566.1
AAI13568.1
AAY14678.1
BAG36187.1
CAA63371.1
CAD44536.1
NP_004473.1
Q14435  
 UDP-GalNAc:polypeptide α-N-acetylgalactosaminyltransferase T6 (GalNAc-T6) 2.4.1.41 Homo sapiens AAH35822.2
AAI14506.1
BAB15297.1
BAC11118.1
BAG52560.1
BAG59418.1
BAJ10977.1
CAA69876.1
CAF85889.1
CAM83953.1
CAM83954.1
NP_009141.1
NP_009141.2
D7UNW5
Q8NCL4
Q8NCL4
 
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase 5 (ppGalNacT5;GALNT5) 2.4.1.41 Homo sapiens AAF15313.1
AAI42677.1
AAP34404.1
ACC22260.1
BAF84843.1
CAB65104.1
CAD44537.1
NP_055383.1
XP_050509.1
A5PKZ1
Q7Z7M9
Q7Z7M9
 
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase 7 (Galnt7) 2.4.1.41 Homo sapiens AAH35303.1
AAH46129.1
AAH47468.1
BAG52155.1
CAB60270.1
CAC39783.1
CAD44538.1
CAI72154.1
NP_059119.1
B3KQU3
Q86SF2
Q86SF2
 
 polypeptide GalNAc transferase 8 (GalNAc-T8) 2.4.1.41 Homo sapiens CAD44539.1
NP_059113.1
Q9NY28  
 polypeptide N-acetylgalactosaminyltransferase 9 (GalNAc-T9) 2.4.1.41 Homo sapiens AAH69354.1
AAH93817.1
AAH93817.2
AAH93819.1
AAH93819.2
AAM49722.1
AAT18857.1
ABE14200.1
BAB13699.2
BAG51429.1
BAG51570.1
CAD44540.1
NP_068580.1
Q9HCQ5
Q9HCQ5
 
 UDP-GalNAc: polypeptide N-acetylgalactosaminyltransferase 10 (GalNT10;ppGalNAcT10;pp-GalNAc-T10) 2.4.1.41 Homo sapiens AAH07224.2
AAH72450.1
AAH98444.1
AAP65372.1
AAP65373.1
AAP65378.1
AAP65379.1
ABA33148.1
ABA33149.1
ABA33154.1
ABA33155.1
BAB14676.1
BAC56890.1
BAG54441.1
CAB46378.1
CAD44532.1
CAE91109.1
NP_060010.2
NP_938080.1
Q86SR1
Q86SR1
2D7I[A]
2D7R[A]
 UDP-GalNAc: polypeptide N-acetylgalactosaminyltransferase 12 (ppGalNAcT12;Galnt12) 2.4.1.41 Homo sapiens AAH13945.1
ABA38425.1
BAB15027.1
BAC07181.1
BAD93346.1
CAC80100.2
CAD44541.1
CAG32962.1
CAI17042.1
NP_078918.3
Q8IXK2
Q8IXK2
 
 UDP-GalNAc: polypeptide α-N-acetylgalactosaminyltransferase T1 (GalNAc-T1;ppGalNAcTase-1;Galnt1) 2.4.1.41 Mus musculus AAB58477.1
AAH56215.1
AAH90962.1
BAE28617.1
NP_038842.2
O08912
O08912
1XHB[A]
 Polypeptide GalNAc transferase T3 (GALNT3) 2.4.1.41 Mus musculus AAB09579.1
AAH43331.1
BAE22551.1
CAM20333.1
CAM21878.1
NP_056551.1
P70419
P70419
 
 Polypeptide GalNAc transferase T4 2.4.1.41 Mus musculus AAB58301.1
AAH57882.1
ABA38430.1
BAC28317.1
BAE28303.1
BAE31844.1
NP_056552.1
O08832  
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase 7 (Galnt7) 2.4.1.41 Mus musculus AAH07484.1
AAH49907.1
AAH52461.1
AAK37549.1
BAC28284.1
BAC29461.1
BAE30388.1
BAE30662.1
BAE31034.1
BAE31754.1
NP_653332.2
Q80VA0
Q80VA0
 
 polypeptide N-acetylgalactosaminyltransferase T2 (Galnt2) 2.4.1.41 Mus musculus AAH07172.1
AAH53063.1
AAH59818.1
AAK37548.1
BAC32790.1
BAE26246.1
BAE30483.1
NP_644678.1
Q6PB93  
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase GalNT11 2.4.1.41 Mus musculus AAH11428.1
AAH21504.1
AAH34185.1
AAH36143.1
AAH36145.1
AAQ06668.1
BAC40096.1
BAE36442.1
CAC79626.1
Q921L8
Q921L8
 
 Polypeptide GalNAc transferase T6 2.4.1.41 Mus musculus C57BL/6J AAI19325.1
AAI19327.1
BAC25984.1
BAC33618.1
BAE32989.1
BAE35316.1
CAB55352.1
NP_766039.1
Q0VE84
Q8C7U7
Q8C7U7
 
 Polypeptide GalNAc transferase T6 2.4.1.41 Rattus norvegicus AAC99426.1
NP_075215.1
Q9R0C5  
 UDP-GalNAc: polypeptide N-acetylgalactosaminyltransferase T1 (GalNAc-T1;ppGalNAcT1) 2.4.1.41 Rattus norvegicus AAC52511.1
NP_077349.1
Q10473  
 N-acetylgalactosaminyltransferase T5 (Galnt5) 2.4.1.41 Rattus norvegicus AAC69708.1
NP_113984.1
O88422  
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase T9 2.4.1.41 Rattus norvegicus AAK54498.1
EDM04502.1
NP_570098.1
Q925R7  
 polypeptide N-acetylgalactosaminyltransferase T1 2.4.1.41 Sus scrofa BAA12800.1 Q29121  
 UDP-GalNAc:polypeptide N-acetylgalactosaminyltransferase T3 (ppGalNAc-T3) 2.4.1.41 Toxoplasma gondii AAO20278.1
AAQ95173.1
Q6YBY0  
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Footnotes

Experimental characterization exclusively covers enzyme activities at present. If you are aware of any missing experimental characterization that should be mentioned here, we encourage you to send your comment and support of the experimental evidence, including an electronic copy or appropriate link to the support material (i.e., article(s), thesis, database, etc), to cazy@afmb.univ-mrs.fr.





Last update: 2013-05-13 © Copyright 1998-2013
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